M Sc Dissertation(WII)
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Item Habitat Use by Goral (Nemorhaedus goral bedfordi) in Majhatal Harsang Wildlife Sanctuary Himachal Pradesh, India(Wildlife Institute of India, Dehradun, 1993) Mishra, Charudutt; Johnsingh, A.J.T.Studied the habitat use pattern of goral (Nemorhaedus goral bedfordi} in Majhatal Harsang Wildlife Sanctuary in the Himalaya to determine its habitat requirements. Of special interest were the roles of forage availability and quality and the antipredator strategy of goral in determining its habitat selection. These factors are of paramount importance in influencing the habitat selection by mountain ungulates. Diet composition of goral in terms of the proportions of graminoids versus browse was determined through pellet analysis. Its escape strategy was determined by direct observation. These results were then related to the actual habitat use patterns, which were determined by obtaining and quantifying locations of goral (n=334) over two seasons, along five monitoring trails. Use of each habitat category was interpreted with respect to its availability. A non-mapping technique was used for determining the availability of each habitat component. I identified nine vegetation types based on physiognomy and floristics. These were: Open Pine Community (OPC), Dense Pine Forest (DPF), Open Oak-Pine Community (OOPC), Dense Oak-Pine Forest (DOPF), Nullah Oak Forest with Low undergrowth (NOFL), Nullah Oak Forest with High undergrowth (NOFH), Euphorbia-Woodfordia-Dodoenia Scrub (EWDS), Open Euphorbia Scrub (OES) and Low Altitude Nullah Forest (LANF). Both forage quality and the antipredator strategy had a profound influence on habitat selection by goral. Goral was a grazer. It was partial to the younger, more nutritive grass phenophases - a prediction that can be made on the basis of its small body size. It preferred open areas with extensive grass cover. Forest cover, along with cliffs, was an important escape area for goral. But it avoided areas with extensive shrub cover. Such areas have little grass. Besides, the shrub cover obstructs visibility and quick movement, and makes the animal vulnerable to predation.Item Social Behaviour and Communication Among Wild Lion-Tailed Macaque (Macaca silenus) in the Indira Ganghi Wildlife Sanctuary, Tamil Nadu(Wildlife Institute of India, Dehradun, 2001) Raghavan, Roopali; Gupta, A.K.; Johnsingh, A.J.T.This study on the social behaviour and communication among wild lion-tailed macaque was carried out between November 2000 and April 2001, on a group of lion-tailed macaques that inhabit the wet evergreen Puthutotam forest fragment. located within the Indira Gandhi Wildlife Sanctuary in the Anaimalai hills, Tamil Nadu.The troop consisted of 16 to 19 animals. For the purpose of this study, social interactions within and between adults and subadults of both sexes were considered. Focal animal sampling and instantaneous scan sampling was thus conducted on only ten identified individuals (four males and six females) in the troop. A complete ethogram describing all behaviour observed during the period of the study was repared. The time-activity budget was calculated for the troop. The predominant behavioural states among all the individuals were Active Forage (mean ± SD of 22.4 ± 0.08%), Active Feed (17.0 ± 0.05%), Sit (16.6 ± 0.04%), Move (13 ± 0.030/0), Passive Forage (6.8 ± 0.02%) and Passive Feed (6.5 ± 0.02%). The adult males and females of the study troop differed in the proportion of time spent by them in the different behavioural states. Autogroom and Allogroom accounted for the principal difference between the sexes. Adult males spent relatively greater time Autogrooming than females, while females allogroomed relatively more. Dominance ranks were calculated independently for males and females in the study troop. Among both males and females, rank position in the dominance hierarchy appeared to be influenced strongly by body size. This is an unusual finding for the females of cercopithecine primate species, implying the existence of an unique individual-based dominance hierarchy among the study females. There was i.e remarkable similarity in the frequency with which dominant and subordinate females displayed virtually all behaviours towards each other in dyadic interactions. This provides support to the prevailing view that lion-tailed macaques indeed display an egalitarian nature. Classification of the gestures and vocalisations were observed to be largely dependent on the dominance rank of the individuals. with certain gestures being characteristic of submission. Males and females showed differential use of the gestures and expressions.Item Density, Biomass and Habitat Occupancy of Ungulates in Bhadra Tiger Reserve, Karnataka(Wildlife Institute of India, Dehradun, 2001) Jathanna, Devcharan; Johnsingh, A.J.T.The distribution and abundance of ungulates prey of tiger were studied in the tropical moist deciduous forest of Bhadra Tiger Reserve, Karnataka from November 2000 to April 2001. The species of interest were chital, sambar, Muntjac and gaur. The line transect method was used to estimate absolute densities of the study species in November 2000. Distribution of the study species was studied using dung as an indicator of occupancy. Logistic regression models was used to examine species presence/absence and habitat parameters.Item Health and Sexual Signals of Male Asian Elephants (Elephas maximus) in Nagarahole National Park, Southern India(Wildlife Institute of India, Dehradun, 1999) Nath, Cheryl; Johnsingh, A.J.T.Male Asian elephants (Elephas maxim us) were studied at Nagarahole National Park, southern India from November 1998 to May 1999. Information was collected on at least 30 identified individuals for investigation of the possible use of musth and tusk length by males as signals of health and genetic fitness to female elephants. This study followed an earlier study by Watve and Sukumar (1997) which showed a significant negative correlation between tusk lengths and parasite loads in male elephants of the Mudumalai Wildlife Sanctuary, southern India. In this project, musth was studied in . addition to tusk length as a possible indicator of good health. It was hypothesised that musth and tusk length in male elephants would be related to better health and body condition and hence could potentially be considered as signals of better genetic quality, within the framework of Zahavi's theory of honest signaling (1975, 1977) and Hamilton and Zuk's theory of heritable true fitness (1982), respectively. The indicators of better health and body condition used were a subjectively scored body condition index (SCI) and a quantitative assessment of intestinal helminth parasite density (parasite load). Individuals were identified by a combination of several physical features, photographic records were obtained for height and tusk length measurement and information was collected on body condition, parasite loads, musth intensity and social interactions. The following interesting results were obtained: 1. Individuals exhibiting signs of early musth had significantly better body conditions and lower parasite loads than individuals exhibiting no signs of musth. 2. Longer tusks were not significantly correlated with better body condition, lower parasite loads or musth in this study.3. Dominance behaviours recorded in dyadic interactions among male elephants indicated that dominance was associated with musth, height of the animal and tusk length in a high percentage of interactions (100%, 89% and 78%, respectively).The significant association between early musth and better body condition as well as between early musth and reduced parasite loads indicates that male elephants may require better health in order to attain musth. Hence musth appears to be a potential honest signal of better than average health, in males which are able to attain this condition. This information may be used by female elephants in choosing mates. Social interactions among males corroborated this speculation by showing that musth and height in male elephants may be important determinants of dominance, which in tum may influence female choice and reproductive success. Limited samples of male-female interactions suggested that females discriminate between different males, and that male-male dominance interactions may influence female choice. However, small sample sizes limited the extent to which these findings could be extrapolated in this and other elephant populations.Item Sexual Segregation in the Nilgiri Tahr (Hemitragus hylocrius)(Wildlife Institute of India, Dehradun, 1995) Madhusudan, M.D; Johnsingh, A.J.T.Studied sexual segregation in the Nilgiri tahr Hemitragus hylocrius from November 1994 through April 1995 at Eravikulam National Park, Kerala, India. This Study covered the post-rut phase of the tahr's annual cycle. The objectives of the study were to examine whether sexual segregation exists in the tahr identify modes of segregation, and compare empirical findings with some theoretical hypotheses advanced to explain sexual segregation. I investigated differences in ,the way the sexes occurred in social units, used space, chose forage and budgeted time. Data on group composition and habitat selection were collected and two trails monitored regularly. Availability of forage was quantified in three habitat strata. Differences in dietary quality were estimated with two faecal indices - crude protein and ash. Data on activity budgets were collected from group scans. Throughout the study period, tahr occurred mostly in female groups, bachelor groups, and 1 to a lesser degree, in mixed groups. They showed marked segregation in the use of habitat types. Male groups used the rolling grasslands at the higher elevations intensively, whereas the female groups were distributed in rocky areas dominated by Plebophylllum kunthianum, and in areas of grass interspersed with cliffs. Male and female groups showed distinct differences in their choice of habitat. Males chose areas of high absolute amounts of graminoid forage, whereas females preferred areas of greater security; these were typically areas high slope angles and percent rockiness. - There were no differences in the levels of crude protein and ash in the pellets of male and female tahr. Differences were also found in the time-activity budgets of the sexes. Although all age sex classes spent equal proportions of time feeding, large males spent a greater proportion of their time resting than either small males or females. Females on the other hand, spent a greater proportion of their time moving than did individuals in the two male classesItem Distribution and movement patterns of the Himalayan Black Bear (Selenarctos thibetanus Cuvier) in Dachigam National Park, Kashmir(Wildlife Institute of India, Dehradun, 1989) Saberwal, Vasant; Johnsingh, A.J.T.This study investigates the distribution patterns of the Himalayan Black Bear (Selenarctos thibetanus) in Dachigam Rational Park, Kashmir. It involved basically two components. (i) looking at the differential usage of different parts of the park by bears over a five month study period. and (ii) the spatio-temporal variation in the distribution and availability of fruit during the study. Dachigam National Park of 141 sq. km ranges in altitude from l800m to 4400m. It is divided into Lower (2& sq km) and Upper (115 sq km) Dachigam. Lower Dachigam (LO) is characterized by a broad valley flanked by steep hillsides. Upper Dchigam (UO) is composed of several gullies separated by fairly steep ridges. The valley vegetation is broad leaved, moist deciduous forest with an unusually high abundance pf fruit species. The slopes on either side are dominated by grass (south facing slopes) or a combination of grass, conifers and shrubs (north facing slopes). The vegetation of Upper Dachigam is mainly composed of coniferous species, with alpine meadows in the higher areas. Transects were walked through Lower Dachigam, considered to have over 90% of the Black Bear population of the park. to record animal locations and signs. especially droppings. 100m segments were marked along these transects and sightings and signs plotted accordingly. These segments were the basis of the habitat mapping exercise. A subjective estimate of the abundance of key fruit species was made for each segment. A phenology study looked at the timing of fruiting of key species. Data analysis shows a good, though not significant correlation between sighting frequencies and fruit availability. However there is strong suggestive evidence to indicate that anilla1 movements are largely Controlled by the spatial distribution and phenological status of five to six key fruit species. The lack of significance is probably due to the small data set. The study indicated the lack of defended territories. Instead, as reported from studies on the American Black Bear (Ursus americanus), there are marked seasonal shifts in the use of the home range. Feeding aggregations of up to 20 bears in one hectare are common. Densities of over 1.5 bears per sq km were recorded in Lower Dachigam at times of high fruit abundance. For two months of the year, May and August, black bears disperse out of the park due to very low food availability. Crop raiding (apple, cherry, and maize) is common at this time. The unusually high abundance levels of fruit in Lower Dachigam (due to oak, walnut, and mulberry plantations, over 70 years old) supports a high density of bears at certain times of the year. However, the lack of sufficient fruit at other times of the year is resulting in increasing levels of conflict with the local human population. Problems of oak and walnut regeneration, and the implications of extremely localized oak distribution are discussed. A more detailed study investigating the problem is suggested.Item Ecological separation of four sympatric carnivores in Keoladeo Ghana National Park, Bharatpur, Rajasthan, India(Wildlife Institute of India, Dehradun, 1989) Mukherjee, Shomita; Johnsingh, A.J.T.The study conducted from 5th May 1989 to 9th October 1989 covering summer and monsoon looked at the ecological separation of four sympatric carnivores in Keoladeo National Park, Bharatpur, Rajasthan. The four carnivores were - Jungle cat (Felis chaus), Fishing cat (Felis viverrina), Jackal (Canis aureus) and otter (Dutra perspicillata). Objectives of the study were to determine differences in dietary composition, habitat occupancy and time of activity as well as to tryout methods for studying sympatric lesser carnivores. Five methods were tried out to obtain these Objectives. Line transects and searches during mornings and nights were used to collect data on prey abundance, location of carnivores and their tine of activity. Scats were collected and analyzed to determine dietary composition and see how they differed among jackal, fishing cat and Jungle cat. Otter spraints were not found. The following conclusions are although some amount of overlap is evident an overall difference in habitat use, time of activity and dietary composition is seen. Jackals and Jungle cats are habitat generalists but dietary specialists. "The fishing cat and otter are habitat as well as dietary specialists . Jungle cat and fishhing cat are largely nocturnal Out the Jackal and otter are active throughout the day. Behavioural observations added to the results obtained from the other methods . 100 methods however were found unsuitable for studying lesser carnivores.