M Sc Dissertation(WII)
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Item Exploring ecological separation between Himalayan blue sheep and Himalayan ibex during winter in Indian Trans-Himalaya(Wildlife Institute of India, Dehradun, 2024) Prakruthi, G.M.; Sathyakumar, S.; Lyngdoh, Salvador; Bhatnagar, YashveerAccording to resource selection theory, two related species with similar ecological requirements compete for the same resources. They can only coexist if they are ecologically separated, either by habitat or food preference. This study explored the resource use and ecological separation of ibex (Capra sibirica himalayanus) and blue sheep (Pseudois nayaur) during the winter in the Spiti Valley, focusing on their segregation in space, habitat use patterns, diet, and activity pattern. The study was conducted in Kibber and Chicham areas (57 km2) of Kibber Wildlife Sanctuary, Himachal Pradesh, India. This study slightly deviates from the findings of a previous study by Namgail (2006), which reveals a high overlap in the habitat use between ibex and blue sheep in the Zanskar mountains. Systematic sampling using transects and trails, vantage points, opportunistic sampling, and camera traps was employed. Habitat and vegetation variables were recorded at random points and animal sightings to understand the patterns of habitat use and food selection. Spatial and temporal overlaps were analysed and micro-histological analysis of faecal samples compared the proportions of monocots and dicots in their diets. The study investigated how ungulates utilize habitats and their dietary habits with the availability of resources in their environment. Results showed substantial overlap in activity patterns, with the Jaccard similarity index indicating less spatial overlap between the species. The generalized linear model and Non-metric dimensional analysis indicated segregation based on their habitat preferences. However, there were no significant differences in their diets, the two species differed in the use of habitat at finer scales. Ibex preferred interspersed rocky slopes and slabs because of high rocky areas (escape terrain) and sparse vegetation, trading food for security. Blue sheep preferred less rocky areas with high vegetation cover like smooth slopes. Ibex preferred areas with more escape routes available than blue sheep. Thus, I conclude that Blue sheep, being more abundant than ibex, occupy distinct regions within the study area, and have a competitive advantage. While ibex are present in smaller numbers and confined to a specific region. Understanding their ecological requirements, especially in harsh winters, is crucial for targeted conservation efforts to ensure the sustainability of these species in a rapidly changing environment. Future research should focus on climate change impacts on habitat use and resource availabilityItem Responses of Wild Ungulates to Anthropogenic Pressure and Perception of Local Communities Towards Conservation of Wild Ungulates in Dampa Tiger Reserve, Mizoram India(Wildlife Institute of India, Dehradun, 2021) Chakama, Joonu; Sathyakumar, S.; Kawlni, Lallaianpuii; Kolipakam, VishnupriyaWith the increasing anthropogenic pressures, conservation of ungulates is ever more crucial as they help in maintaining the forest structure. The impact of anthropogenic pressures such as hunting of wild ungulates by local communities in Mizoram, India, is limited and lacks reliable information. This study was conducted in Dampa Tiger Reserve (DTR), Mamit district, Mizoram from January-Mid May, 2021 to quantify the responses of wild ungulates to the anthropogenic pressures, as a measure of conservation efforts. This study focussed on the population and distribution of ungulates in Dampa Tiger Reserve (DTR), Mizoram and assessed how the ungulates responded to human use areas. Stratified systematic sampling was done in an intensive study area (40 km2) to assess the ungulate density, distribution and activity pattern. In addition to that, the perception and attitude of local communities towards wild ungulates were also assessed. Estimates of the distribution, population and activity pattern of ungulates were obtained using single season occupancy model and cameratrap based distance sampling method whereas the local people perception and attitude were analysed based on answers from the semi-structured interview. The study was carried out on the Teirei and Phuldungsei range of the DTR. A total of 40 grids (1 km2 each) were sampled following the above framework. The grids were sampled using Camera-based distance sampling for population estimation in which the cameras were placed at the centroid of the grid. The density of the ungulates in two areas (high and low human use) could not be compared as the anthropogenic disturbance was homogenous throughout the study area. However, the pooled density of wild pig (0.55±0.23) was the highest among the other species, followed by barking deer (0.28±0.06) and sambar (0.20±0.09). The activities of the ungulates were at the peak from the crepuscular period to early morning and late evening. Occupancy pattern showed that barking deer distribution was influenced by the food availability/ground cover including areas near human habitations whereas sambar occurrence was negatively influenced by non-forest and presence of anthropogenic disturbance. Perception survey revealed that villagers showed positive perception towards ungulate conservation, however, in contrast to these responses, conservation practices in DTR by the local communities still lacks as intensive hunting still prevails. This study highlights the impact of anthropogenic pressure towards ungulates and how strengthen management interventions is necessary to ensure the conservation of ungulates in DTR.Item Playing it Safe Tiger Movement and Livestock Depredation Patterns in Panna Tiger Reserve, Central India(Wildlife Institute of India, Dehradun, 2021) Mohan, Manu K.; Ramesh, K.; Sathyakumar, S.The movement of large carnivores in human-dominated landscapes is not uncommon anymore. While their presence is superficially tolerated or ignored, it is a consistent cause of concern to the people living in its proximity, due to competing interests in sharing of resources and direct threat to human lives. Carnivores incur energetic costs in the form of restricted activity and having to make spatial and temporal activity adjustments. Humans, on the other hand, face losses from livestock depredation and/or loss of lives. Depredation patterns of livestock by carnivores are associated with a variety of factors including habitat structure as well as movement patterns of carnivores in relation to anthropogenic and livestock activity. Hence, deducing movement patterns of the groups in conflict is an important step in understanding the dynamics of conflict patterns at fine scales. Supplemented by models to predict the probability of conflicts spatially which delineates conflict hotspots, site-specific mitigation measures could be devised. To understand such patterns of carnivore movement and relation to livestock depredation events, I conducted this study in a human-dominated region adjacent to the Critical Tiger Habitat of Panna Tiger Reserve in central India from December 2020 to April 2021, within an area of 200 km2. The broad objective was to model the conflict probability based on the proportion of the human-dominated area used by tigers, its intensity of use and spatio-temporal activity within the area, in response to human disturbances. For this, I used intensive camera trap sampling at fine-scale within grids (1 km x 1 km) across the entire study area. Movement parameters of tigers such as displacement and distance to villages in day and night were also analysed by monitoring two GPS-collared tigers in the study area. Occupancy estimates showed 74% of the sampled area being used by tigers, with the relative abundance of a large bodied wild prey (sambar) as the strongest predictor. In addition, a clear positive trend in the relative abundance index of tigers with distance to villages was observed. The temporal activities of the tigers also showed a significant difference in median activity time, shifting towards the night along with moderate diurnal activity at higher distances from villages. The tigers exhibited both crepuscular and nocturnal activity at close distances to villages. The movement range analyses of the tigers using camera traps showed the study area being used by 17 tigers, with 9 of them being adults (>3 years of age). Fixed Kernel Density home-range estimates of the two GPS-collared tigers showed a considerable proportion of their home range areas falling within village boundaries (10%) and an average of 16% of all GPS locations of the tigers fell within 0-100 metres from villages. Despite such high exploratory use of areas within and close distances to villages by sub-adults, there were relatively low incidences of conflict between the tigers and villagers in the study area. Finally, to assess the extent of risk and predict the probability of livestock depredation in the area, Generalized Linear Models were used. The best-fit model indicated a significant negative effect of distance to villages and highway to the probability of livestock depredation. Linear intrusions through highway and forest trails from villages allow easy access to livestock and herders for the utilization of the multi-use buffer areas of the Reserve as well as the tigers to come in proximity to settlements. Corralling of livestock in the night, avoiding free-ranging and grazing of livestock in the identified conflict hotspots, incentivization of alternate source of livelihood by strengthening the institutions such as Eco-Development Committees, along with continuous monitoring of tigers which range close to human settlements are discussed as some of the important measures which can be adopted to decrease the interfaces leading to tiger-human conflict in Panna Tiger Reserve.Item Comparative Analysis of Abundance Estimation Methods for Ungulates Along with their Spatio-Temporal Interaction and Habitat use in Pakke Tiger Reserve, Anunachal Pradesh(Wildlife Institute of India, Dehradun, 2021) Singh, Tarun; Kolipakam, Vishnupriya; Qureshi, Qamar; Sathyakumar, S.Abundance estimation and habitat utilization of species are critical for the conservation of any species. The forests of North-Eastern India are challenging due to the low abundance of ungulates, dense undergrowth, and logistic feasibility. In this study, we attempt to use camera trap-based methods (distance sampling and random encounter model) to estimate the abundance and habitat use of five ungulate species in the Pakke Tiger Reserve of western Arunachal Pradesh. This study was conducted from February 2021 to May 2021.This study provides crucial information on abundances and habitat use, which can be used for focussed conservation efforts for ungulates occurring in low densities.Item Habitat use by Gaur (Bos gaurus gaurus) and Assessment of Human-Gaur Conflicts Human Modified Landscapes of Nilgiris Forest Division Tamil Nadu(Wildlife Institute of India, Dehradun, 2019) Indira, S.; Sathyakumar, S.Gaur is the largest bovine in the world and is threatened by factors such as habitat loss, poaching, diseases and other anthropogenic stresses. In unprotected landscape, their habitat is confined to small forest pockets which are interspersed with human habitations. This study investigated the use of human-modified landscapes by gaur and how does this potentially increase the negative interaction between human and the animal. Information on ecology of gaur is limited to a few studies conducted within protected areas in India. This study was conducted in the six ranges of the Nilgiris FD, viz., Udhagai North, Udhagai South, Kattabettu, Kotagiri, Coonoor, and Kundha that encompassed an area of 900 km2 during the period January to April, 2019. Secondary data related to conflict and gaur mortality was collected from the District Forest Office to assess the current conflict scenario. To investigate the habitat use by gaur, the study area was divided into grids (2 km x 2km) and an average effort of 1.5 km (+/- 0.5 km) was made in every grid. To evaluate the perception and attitude of people towards gaur and human-gaur conflict, a semi-structured questionnaire was designed preceding the reconnaissance survey. The total crop raiding incidences were 31, human deaths 6, human injury 36, livestock death 4 and property damage 3. Injuries to humans due to gaur attacks appears to be most recorded conflict as indicated by the number of incidents in 3 years, followed by crop raiding, human deaths, livestock deaths and property damage. Areas such as Nunthala, Adhigaratti and Ithalar of Kundha and Udhagai south has high conflict. Areas such as Konvakarai and Bickapathy had medium conflicts and the rest were of low conflict intensity. Most of the conflicts were of crop raiding during the months January to April. In total, 79 gaur sightings 38 dung were recorded in the study area during the study period. Most of the visual sightings were in tea plantation followed by reserve forest, human settlement and cropland. Gaur dung were recorded only in forested habitats. Various groups of people majorly sight gaur in tea plantations than other habitats. Most of the respondents have sighted gaur in and around human habitation during dawn followed by dusk and night hours. Crop Raiding (CR), Human Attack (HA), Traffic hindrance (TRA), Presence in Human Habitation (PRE HH) are the classes of conflicts perceived by the local communities. Majority of the respondents sighted gaur at least twice in a weak. Respondents’ perceived gaur as threat majorly due to have experiencing fear of attack and crop raiding. Gaur is considered as a major threat animal followed by sloth bear, wild pig, leopard and bonnet macaque. Tea plantations are the source for grass and weeds for the gaur to feed on. Proportion of reserve forest is less than other land use classes in the study area. Hence, for the large mammal like gaur can sustain only if it adapts to use the human habitation for food. Investigations on body health condition, home range, population, breeding ecology, behavior, and so on should be made in the future to understand the ecology of the large bovid within close proximity to human. Such research findings will help to make better conservation efforts for site specific and species specific issue. In spite of various wildlife encounters and conflict incidences it is likely that people and animals are co-existing in the Nilgiris. The Forest Department may have to adopt proactive measures for monitoring and management of human-animal conflict in the Nilgiris with more emphasis on gaur. Awareness creation amongst local communities and rapid response by the Forest Department personnel to manage human-wildlife conflict is crucial to avoid injury/ death and other losses/disturbances for the peaceful co-existence of both animal and people.Item Response of wild mammalian species to human-mediated resource base in Chamoli district of Uttarakhand India(Wildlife Institute of India, Dehradun, 2017) Shrivastava, Nimisha; Sathyakumar, S.; Ramesh, K.This study focuses on the wild animals that occur in human-modified landscapes, in line with the proposed national strategy to manage wildlife in such landscapes. With the aim of focusing on the factors that drive or facilitate species to thrive in human modified areas, I carried out an investigation on the ‘response of wild animals to human modified landscape in Mandal Sub-basin (ca 140 km2), Chamoli district, Uttarakhand’ from December 2016 to April 2017. 2. For the study, I assessed distribution and use of various land use types (human inhabitations, crop fields, secondary scrub around villages, moist and dry natural forest/habitats) by wild mammals using camera traps deployed within grids (1 km2) for a period 10-15 days. Local peoples’ perception surveys were also carried out. Major findings of the study includes a) there are species that are largely dependent on modified landscapes while others that are dependent mostly on natural habitats, b) there appears to be an adaptive strategy to avoid human interaction by adjusting activities in different time scale between natural forests and human dominated areas, c) factors that are responsible for occurrence of these species in various land use types, and hence getting an idea of what future holds for these animals and d) positive perception of local people that shows a light of hope on the conservation of animals in their lands. However, it cannot be done if locals do not get direct benefits by the presence of wild animals.Item Ecology of Leopard (Panthera pardus) in Sanjay Gandhi National Park Maharashtra with Special Reference to its Abundance Prey Selection and Food Habits(Wildlife Institute of India, Dehradun, 2015) Surve, Nikit; Sathyakumar, S.; Sankar, K.Though information is available on the leopard's ecology from natural habitats, very less information is available on the ecology of leopards from human-dominated habitats in India. Hence the study was conducted at Sanjay Gandhi National Park (SGNP), Mumbai where leopard was found to be the apex predator. The total area of SGNP is 104 sq.km. It is covered , by human habitations on all the sides leaving just a narrow tip on the northern side. Camera traps were deployed in a systematic manner covering the entire study area to get leopard images. Leopard density of 21.55 ± 4.6 (SE) at 95% confidence interval was calculated in the study area using Spatially Explicit Capture Recapture models. The wild prey densities were estimated inside SGNP using 'Distance sampling with the help of line transects. The , maximum density amongst ungulates was shown by chital (14.49 ± 5.2[SE]) followed by sambar (7.52 ± 1.7[SE]). Among primates, maximum density was shown by common langur (48.04 ± 8[SE]) followed by bonnet macaque (30.78 ± 12[SE]). Density for other prey species encountered on line transects like wild pig and barking deer were not estimated as they were sighted only twice on the line transects. Dog density on the periphery of SGNP was ca1culated to be 17.26 ± 0.69 (SE) using mark-recapture technique. Food habits of leopards were studied by conducting dietary analysis using leopard scat. The high density of dogs in the periphery of the National Park also reflected in the leopard's diet. Dogs alone contributed 24.46 % of the total biomass consumed in leopard's diet. Domestic prey contributed to 43% whereas wild prey contributed to 57 % of leopard's diet in terms of relative biomass consumed. The study suggests that the leopards are dependent on both wild as well as domestic prey in the study area. Leopards can coexist with humans even amidst urban environs as long as prey population (both domestic and wild) remain sufficient and their habitat is protected.Item Nutritional Ecology of Asian Elephants (Elephas maximus) in Chilla Range of Rajaji National Park, Uttarakhand(Wildlife Institute of India, Dehradun, 2009) Datta, Suniti Bhushan; Goyal, S.P.; Sathyakumar, S.Responses of animals to the habitat has been the central focus for management of species and of these aspects, nutritional ecology has been the key issue in foraging behaviour studies and habitat studies. Therefore, the study focused on the nutritional aspect of forage selection by Asian elephants (Elephas maximus) in Chilla Range of Rajaji National Park. The main objectives were to determine whether habitat structural heterogeneity or nutritional quality of the selected food plant species was a determinant in the temporal and spatial habitat use by elephants and to determine the reasons behind seasonality in foraging. The study area of 148km2 in Chilla Range of Rajaji National Park was divided into twelve 2x2km grids, and these were further divided into 4 sub-grids each. In each sub-grid, a 1 km long line transect was laid and along this 10m radial plots were placed at 100m intervals. Vegetation data were collected to characterize the habitat structural heterogeneity in terms of number of trees of each species, height of canopy base, canopy volume, percent canopy cover, species diversity, species density and geo-spatial variables such as NDVI, standard deviation of slope, mean elevation and the proximity to water. Parts of 12 most-preferred plant species were collected in the study areas for estimating nitrogen, ash, acid detergent fibre, and macro and micro minerals such as sodium, potassium, calcium, copper, magnesium and zinc. During the study period from December 2008 to May 2009, Elephant response was determined in terms of dung density along the 1km transect in winter and in summer. The total number of trees in each grid varied from 124 to 268. Tree diversity ravged betweel) 0.456-1.454. Height of canopy base was mostly 1.5-3.0 m, although two grids showed extremely high canopy bases. Canopy volume ranged from 165.63m3 to 948.36m3 , although the majority of the grids showed a high variation in canopy volume in terms of standard deviation (SE). The percent canopy cover ranged from 54.13% to 93.72%. Most of the NDVI values were _high in the study area (>0.180), while one grid showed a low value of 0.143. For the standard deviation of slope, the values ranged from a low of 2.46 to a high of 8.46. The mean elevation of the study area ranged froin 391-840m a.s.l. Nutritional values estimated for most of the parameters in winter and summer from selected plant samples collected in the study area showed slight variations 4 between seasons. Percent nitrogen content indicated no seasonal difference (f=0.98, p=0.05, df=ll). The percent acid detergent fibre indicated a difference (P<0.05,df=I I) between the two seasons. Percent ash content was found not significantly different (p=O.l6, p=0.05, df=l1) between seasons. Amongst the macro and micro minerals, sodium content was significantly different across the two seasons. Potassium content was found to be significantly different across the seasons (p<0.05). Calcium content was also found to be significantly different between winter and summer (P<0.05). The copper content could not be compared across the seasons as in summer the levels present in plant samples were too low to be detected by the instrument. The difference in magnesium and zinc content across the two seasons were not significant (p=0.16 and P=0.31). Dung density in the grids across both seasons was non-uniform and highly skewed (p<0.05, df=47). The dung densities in both seasons were related more with the habitat heterogeneity variables than nutritional values obtained in each grid. The relationship between dung density and the number of plants was positive in the winter seasons (R2= 0.2848) and summer (R2= 0.4383), indicating that elephants are highly selective towards areas with higher numbers of woody plants. Plant species diversity indicated no influence on dung density in winter (R2= 0.00005), but showed a negative trend in summer (R2= 0.0154). The height of canopy base was negatively related to dung density, with elephants selecting areas with a mean canopy base height between 1.5-3.0m during both winter (R2=0.2288) and summer (R2=0.174). Dung density showed a negative trend when related to canopy volume in both seasons. The R2 value for winter is 0.2087 and in summer it is 0.1471. Percent canopy cover had a negative influence on the dung densities in winter (R~= 0.083) and in summer (R2= 0.1524). NDVI showed a negative relationship with dung densities in winter (R2=0.01l1) and a positive relationship in summer (R2= 0.1894). The relationship between the standard deviation of slope and dung density showed a negative trend in both winter, (R2=0.0033) as well as in summer (R2=0.0389). The higher elevation grids show a lower d/mg density during both winter (R2=0.216) and summer (R2=O.l443). The relationship between dung density and proximity to water in winter is negative (R2=0.1575) and the relationship remains negative (R2=0.1016) in summer. In relation to nitrogen content and dung density in winter there was a weak positive trend (R2=0.0256), while in summer there was a weak negative trend (R2=0.032). The relationship between dung densities and percent acid detergent fibre (ADF) in winter indicated a weak positive trend (R2=0.0012) and during summer, indicated a weak negative trend (R2=0.0657). In relation to percentage ash content (Fig. 4.29.), dung densities in winter indicated a weak positive trend" (R2=0.0114), while in summer there was a weak negative trend (R2=0.0641). When compared with sodium, dung densities showed a very weak positive trend (R2=0.0092) in winter and a negative trend in summer (R2=0.0834). In winter, when compared with dung densities potassium showed no trend (R2=0.001), but in summer, there was a weak negative trend (R2=0.0076). In winter, calcium does not show any relationship with dung densities (R2=0.0002), while in summer, there is a weak negative trend (R2= 0.0511). Magnesium content does not show any relationship with dung densities (Fig 4.33) in the winter season, (R2=0.0007), while in summer, there is a weak negative trend (R2=0.004). Zinc shows a weak positive trend (R2=0.0298) in winter and a weak negative trend (R2=0.026) in summer. Principal component analysis of nutritional parameters indicated nine components that were influencing dung density distribution in the study area in both the seasons, and hence no single parameter influenced elephant habitat use. The study clearly indicates that the distribution of elephants in Chilla Range of Rajaji National Park is more related to abundance of woody species and proximity of water, than the nutritional content in plant species. Principal component analysis showed that there was no single parameter that influenced dung distribution in the study area. This could be due to the fact that plant species selected for foraging by elephants contained an adequate amount of nutrients in most species for meeting foraging requirements .an d they probably meet their daily requirements by foraging on varied proportions of plant species.Item Assessment of Human-Elephant Conflicts in Jalpaiguri District of West Bengal in Relation to Land-Use Patterns(Wildlife Institute of India, Dehradun, 2003) Roy, Anirban Datta; Goyal, S.P.; Sathyakumar, S.This study on Human Elephant Conflict was conducted in Jalpaiguri district .of north West Bengal to identify the level of conflicts in different areas of the district, and to recognize it in connection with land-use patterns existing in the district. The study was conducted over five months and information on conflicts was collected from Tea Gardens and Wildlife Squads. Interviews and questionnaires were also used to survey the conflict affected viilages. The different conflict areas and the intensity of the conflicts in those areas were identified with the help of gridded maps. Results showed that although most of the conflict areas had remained the same from previous studies, some new conflict areas were also emerging. Seasonality of the conflicts was also found to be related to cropping patterns adopted. Information collected from the Tea gardens showed that the Tea gardens which had previously been categorized as high conflict areas had remained same to a great extent. The Tea gardens having the highest conflict incidences were frequented more by solitary animals rather than herds. The mean group size of the herds were found to show two distinct peaks in the months of May-June and September-November. Interviews with affected villagers provided a true picture of the difficulties faced by them during crop-raiding. In the absence of Wildlife Squads, villagers had to drive the elephants away with their own methods.Item Food selection and ranging in hoolock gibbon (Hylobates hoolock Harlan 1834) in Borajan reserved forest, Assam(Wildlife Institute of India, Dehradun, 1997) Kakati, Kashmira; Sathyakumar, S.This study on food selection and ranging in the hoolock gibbon (Hylobates hoolock ,Harlan 1834) was carried out between December 1996 and April 1997 in Borajan Reserve Forest, a disturbed and fragmented patch of remnant tropical evergreen forest in Upper Assam, India. The hoolock gibbon is a primarily frugivorous species, confined to closed-canopy evergreen forests of North-east India, Bangladesh and Burma. This type is highly endangered in its entire range. Threats to the continued existence of the hoolock are from large-scale destruction of its habitat and hunting. Two troops of gibbons were followed intensively for four months and behavioural data were collected by continuous focal animal observations. Data were also collected ad libitum on a third troop. Data on the structural components of the vegetation and on the availability of food resources in the Intensive study area were collected systematically. The results of the study show that food availability influenced the diet, movement patterns, home-range sizes and behaviour of hoolock gibbons. Gibbons were selective in their diet using only forty-three plant species out of the more than two-hundred species present. Figs constituted important keystone resources for the gibbons. In months of low fruit availability, the gibbons had a predominantly folivorous diet. Home ranges were small (10.4 ha and 5.4 ha) and habitat degradation represents the greatest threat to the gibbons