M Sc Dissertation(WII)
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Item Interaction of Kiang (Equus kiang) with Livestok in Hanley Valley of Chanthang Wildlife Sanctuary, Ladakh(Wildlife Institute of India, Dehradun, 2009) Hussain, Asif; Qureshi, Qamar; Rawat, G.S.Hanley Valley «320 41' 27" N 790 04' 3.5") in Eastern Ladakh forms the western most extension of Tibetan plateau and has been recognized as an important biogeographic province in India (Indian Trans-Himalaya). This region harbors a rich array of wild and domestic ungulates. The area is also home to nomadic Changpa and Tibetan refugee herders. The political, social and ecological transformations have altered previous, well-established links between the pastoral population and their rangeland environment. Over the years, with growing integration of the local economy of Ladakh and with better development of cash markets, the pastoral community is fast losing its tolerance towards the kiang and it is increasingly seen as a competitor to livestock. The purpose of this study was to document this intensifying conflict between pastoralism and Kiang conservation, using resource selection functions. Data on habitat variables were collected on 4 fixed trails of varying length (3- 7 km) for kiang (with and without livestock presence) and livestock for use availability analysis. For food habits, micro-histological method in case of kiang and bite count method in case of livestock was used. Data on activity pattern of kiang was collected to examine change in response to livestock presence. Density and encounter rates were estimated using vehicle transects (n=5) of varying lengths (5-56 km). Livestock were followed (focal animal sampling) from dawn to dusk to estimate amount of forage removed. Availability of different habitat variables was estimated using ArcGIS 9.2. Total of 104 kiang groups before arrival of livestock and 187 kiang groups after livestock arrival to the area were recorded. Habitat use by kiang and livestock was significantly different (8 = 0.00016, P < 0.05). Habitat use by kiang before and after livestock arrival was found to be significantly different (8 = 0.013, P <0.05). Groups of kiang were observed (n=14) for time budget evaluation, 7 prior (640 min) and 7 (820 min) after livestock had come to the area. Activity pattern of kiang before and after livestock arrival was found to be significantly different (8 = 0.04, P <0.05). Food preference for kiang before and after livestock arrival was not found to be significantly different. Food preference for kiang and livestock was significantly different (6= 0.0000007, P < 0.05). Use-availability analysis suggested differential selection for food plants. Schoener's niche overlap for food items found moderate overlap between kiang, sheep and goat «60%) and high overlap between kiang and horse (>80%). Forage removed (kg) by sheep, goat and horse were 2.33 kg, 2.25 and 8.19 kg respectively. Total density and encounter rate for Hanley valley was 0.07 kiang/km2 and 0.45 kiang per km respectively. The difference in habitat use translated into Kiang using higher elevations while livestock used middle elevations more than available. Kiang used steep slopes whereas livestock used steep as well as gentle slopes more than available. Kiang used North-West aspect more than available. Kiang used vegetation communities dominated by graminoides while sheep-goat used communities dominated by shrub and forbs. Density and encounter estimates did not show considerable increase in kiang population compared to previous studies. Hence, the perception of people of kiang overstocking in the study area appears to be misplaced. Detailed perception studies should be carried out in the whole of Changthang and herder-centered participatory programmes need to be carried out on a large scale to ensure long term conservation of kiang in eastern Ladakh.Item A Study of Vigilance Behaviour of Chital (Axis axis) in Pench Tiger Reserve, Madhya Pradesh(Wildlife Institute of India, Dehradun, 2009) Ghuman, Sartaj S.; Sankar, K.Predation is an important agent of natural selection in animal communities and so vigilance behaviour has very high selection pressure acting upon it. During feeding bouts, unless plant density and biomass are high and food intake is limited only by the processing rates, herbivores tend to experience a trade-off between vigilance and feeding. Thus, vigilance as a behaviour pattern ought to be tightly regulated by the cost-benefit ratios. A study on vigilance behaviour of chital (Axis axis) was conducted at Pench Tiger Reserve, Madhya Pradesh from December 2008 to April 2009. The objectives of the study were to examine the determinants of individual and group vigilance in chital; and also to examine the t effect of conspecific behaviour on individual vigilance. I used scan sampling with fixed interval recording to estimate group vigilance and focal animal sampling with continuous recording to estimate individual vigilance levels. Three main factors that are known to affect :, vigilance from studies on other herbivores were examined: group size, density of animals in the group and visibility. Scanning behaviour was used as an estimate of vigilance. 'Scanning' refers to the state when the animal has its head above shoulder level and the senses of sight, smell and hearing are actively 'scanning' the environment. Individual vigilance was found to decrease with increasing group size, decreasing density and increasing visibility. In large groups adult males, fawns and adult females with fawns were found to be significantly more vigilant than females without fawns and yearling males. Group vigilance level was found to increase significantly with increasing group size, even as the group scan level decreased. Individuals seemed to reduce vigilance with relation to group size rather than the group vigilance level. The results suggest that individual chital do not adopt levels of scanning that maximize cooperative predator detection. The benefits of the group size effect vary with age-sex classes and other probable internal and external predation risk Factors and so vigilance is more likely governed by a combination of dilution and detection hypotheses mechanisms, the focus changing from one to the other with individual risk.Item Habitat use and food selection by wild and domestic ungulates in the Sikkim Transhimalaya.(Wildlife Institute of India, Dehradun, 2007) Chanchani, Pranav; Rawat, G.S.By defining a resource, determining the resources available to animals and sampling the array of resources actually used by an animal (Krebs 1999), it becomes possible to gauge the nature of interactions between species. This study explored aspects of resource use by diverse assemblage of wild and domestic herbivores including The Tibetan argali (Ovis ammon hodgsoni), Tibetan Gazelle (Procapra picticaudata), kiang (Equus kiang), blue sheep (Pseudois nayaur), domestic yak and sheep in a Trans-Himalayan environment during the lean winter period. Sampling was carried out in a systematic manner using trails, as well as by sampling opportunistically. To quantify vegetation, a 3.52 Ian grid was overlaid on an image of the study area, and grids were randomly picked from these for random sampling. A number of habitat and vegetation variables were measured or noted for all ungulate sightings or within vegetation sampling stations and these were used in analysis to ascertain patterns of habitat use and food selection. U sing a hierarchy of spatial scales, the study modeled animal distributions, and investigated the use of habitats and food habits of ungulates in relation to the availability of these resources in their environment. The study's major findings were that all ungulates were found to occur in the true Trans-Himalaya except for the blue sheep which selected more mountainous terrain in the transition zone between the greater and Trans-Himalaya .. Statistical tests of significance revealed that ungulates differed from one another or from random in their use of resources. Further, terrain features appeared to influence habitat selection to a greater-extent than vegetation. Finally, the findings of this research indicate that although the wild and domestic ungulates of this region all exist in relatively large numbers, they tend to vary in their use of habitats and food either by differences in their distribution, or in the selection of finer environmental (habitat and food) variables. The study therefore concludes that competitive inter-specific interactions are not very apparent in this region.Item Habitat use by Chital (Cervus axis) in Dhaulkhand, Rajaji National Park, India(Wildlife Institute of India, Dehradun, 1993) Bhat, Sridhar D.; Rawat, G.S.A study on the habitat use by chital or spotted deer (Cervus axis) was conducted in Dhaulkhand Rajaji National Park, from November 1992 to May 1993. The objectives of the study were to understand the spatio-temporal use of habitats by chital and to identify the factors that govern the patterns of habitat use. Foot transects were used to estimate the densities of chital and quantify the availability and utilization of resources.Item Habitat Use by Goral (Nemorhaedus goral bedfordi) in Majhatal Harsang Wildlife Sanctuary Himachal Pradesh, India(Wildlife Institute of India, Dehradun, 1993) Mishra, Charudutt; Johnsingh, A.J.T.Studied the habitat use pattern of goral (Nemorhaedus goral bedfordi} in Majhatal Harsang Wildlife Sanctuary in the Himalaya to determine its habitat requirements. Of special interest were the roles of forage availability and quality and the antipredator strategy of goral in determining its habitat selection. These factors are of paramount importance in influencing the habitat selection by mountain ungulates. Diet composition of goral in terms of the proportions of graminoids versus browse was determined through pellet analysis. Its escape strategy was determined by direct observation. These results were then related to the actual habitat use patterns, which were determined by obtaining and quantifying locations of goral (n=334) over two seasons, along five monitoring trails. Use of each habitat category was interpreted with respect to its availability. A non-mapping technique was used for determining the availability of each habitat component. I identified nine vegetation types based on physiognomy and floristics. These were: Open Pine Community (OPC), Dense Pine Forest (DPF), Open Oak-Pine Community (OOPC), Dense Oak-Pine Forest (DOPF), Nullah Oak Forest with Low undergrowth (NOFL), Nullah Oak Forest with High undergrowth (NOFH), Euphorbia-Woodfordia-Dodoenia Scrub (EWDS), Open Euphorbia Scrub (OES) and Low Altitude Nullah Forest (LANF). Both forage quality and the antipredator strategy had a profound influence on habitat selection by goral. Goral was a grazer. It was partial to the younger, more nutritive grass phenophases - a prediction that can be made on the basis of its small body size. It preferred open areas with extensive grass cover. Forest cover, along with cliffs, was an important escape area for goral. But it avoided areas with extensive shrub cover. Such areas have little grass. Besides, the shrub cover obstructs visibility and quick movement, and makes the animal vulnerable to predation.Item Habitat use by the Great Indian Rhinoceros (Rhinoceros unicornis) and the other sympatric large herbivores in Kaziranga National Park(Wildlife Institute of India, Dehradun, 2001) Banerjee, Gitanjali; Rawat, G.S.; Choudhury, B.C.A study on the habitat use by the Great One horned rhinoceros (Rhinoceros unicornis) and three other sympatric ungulates was carried out in Kaziranga National Park (KNP), Assam during December 2000 to April 2001. KNP supports more than half the world's population of the Indian Rhinoceros, a highly endangered species. Since this park also supports a high diversity and density of herbivores coexisting in a typical floodplain habitat, it was an ideal place to conduct the following study. The study aimed to determine how 4 large herbivore species rhino, wild buffalo, swamp deer and hog deer achieve spatial and temporal separation across two seasons. The study also investigated how nutritive content of the forage determines habitat selection by these species. KNP was an ideal place to study habitat use patterns by these four sympatric species due to the seasonal variation of forage quality caused by the practice of annual burning. Sampling for animal abundance within each habitat type over two seasons was done by monitoring transects. Feeding observations were obtained by scan sampling. Laboratory analysis was done to determine the nutritive content of the available forage during winter and summer. Seasonal and spatial differences observed in habitat occupancy patterns by the ungulate species within KNP seem to be a way of partitioning resources in order to minimize competition. During the winter season all ungulates selected the short grasslands for feeding. The wild buffalo and the swamp deer showed no variation in the habitat occupancy patterns showing a positive selection for the short grasslands. However, rhinos and hog deer were observed to prefer the tall, burnt and sprouting grasslands in summer for feeding during summer. When habitat occupancy patterns were correlated with the nutritive value of the forage it was found that all ungulates tracked high crude protein levels in available forage. Rhinos and hog deer formed an association that exploited areas that have high crude protein and low silica content. Wild buffaloes and swamp deer show less flexibility in habitat occupancy patterns and were observed to feed in short grasslands where there was a relatively high amount of crude protein available during both the seasons. The study revealed that crude protein played an important role in determining habitat use by hog deer, which shows a linear relationship with crude protein. Forage volume, an indicator of available food played a significant role in determining habitat use patterns for the large bodied ungulates, namely the rhino and the wild buffalo which is in keeping with their physiology and body requirements. There seems to be partitioning of space by the ungulates on the basis of body size and dietary separation. Rhino and hog deer were observed to formed one guild whereas wild buffalo and swamp deer formed another one. These two groups differed in the pattern of habitat occupancy suggesting that ecological separation in Kaziranga National Park, perhaps occurs on the basis of differential use of the habitat by ungulates that have a similar body size. There was an observed complementation of body sizes within each association. A large bodied ungulate associated with another ungulate that was much smaller in size. and therefore, had different ecological needs.Item Resource selection and resource partitioning among wild ungulates in the tropical semi-arid forest of Ranthambhore National Park. Rajasthan.(Wildlife Institute of India, Dehradun, 2001) Bagchi, Sumanta; Goyal, S.P.; Sankar, K.This study attempted to explore and describe the pattern of resource selection and inter-relationship in communal niche space of four species i.e. chital, sambar, nilgai and chinkara in dry deciduous semi arid forests of Ranthambhore national park, RajasthanItem Ranging, Activity Patterns and Habitat Use of Blackbuck and Nilgai in Velavadar National Park, Gujarat, India(Wildlife Institute of India, Dehradun, 2001) Sahabandhu, H. Dhanushki R.; Chellam, Ravi; Mathur, V.B.I studied the activity patterns, ranging and habitat use of blackbuck (Antelope cervicapra rajputanae) and nilgai (Boselaphus tragocamelus) in Velavadar National Park from November 2000 to April 2001. Scan sampling was used to record blackbuck and nilgai activity patterns. Herds were also followed from sunrise to sunset to study the diurnal movements. Satellite imagery was used for vegetation mapping. Habitat availability and usage of both species were examined by plotting the grazing circuit on the satellite imagery. Activity patterns were found to differ amongst the various social groups, habitats and seasons. Temporal variations in foraging time and resting time were found in all the groups studied. Blackbuck had three foraging peaks and two resting peaks as compared to nilgai, which had two, and one peaks respectively. Diurnal distances were not found to differ in any of the different blackbuck or nilgai groups or between the two species. Differences between blackbuck herds in different seasons and habitats were attributed to the nutritional levels and the spatial dispersion of vegetation. Foraging behaviour decreased in summer supporting results of previous studies that blackbuck reduced foraging due to lower nutrition levels. Male blackbuck was found to forage 67% compared to 58% by females. This difference was attributed to larger body size in males, and the upcoming rutting/lekking season. No difference was found between nilgai males and females, except usage by females was slightly more in high nutritional areas. Female nilgai were found to become more selective in summer, by increasing their foraging time from 43% to 55%. But no difference in habitat usage was found, possibly due to sustenance from Prosopis juliflora pods. Blackbuck were found to spend more time foraging (53%) compared to nilgai (43%). This was due to blackbuck being more selective in the prevailing drought conditions, while nilgai were possibly supplementing their diet with P Juliflora pods. Differences between the two species in temporal allocation of foraging time was found which was ascribed to difference in gut capacity. No seasonal differences were found between the two species, except that nilgai were found to be using high nutrition areas. Nilgai were found to be more selective nutritionally than blackbuck, possibly due to nilgai being an intermediate feeder compared to blackbuck, which are coarse bulk feeders.Item Density, Biomass and Habitat Occupancy of Ungulates in Bhadra Tiger Reserve, Karnataka(Wildlife Institute of India, Dehradun, 2001) Jathanna, Devcharan; Johnsingh, A.J.T.The distribution and abundance of ungulates prey of tiger were studied in the tropical moist deciduous forest of Bhadra Tiger Reserve, Karnataka from November 2000 to April 2001. The species of interest were chital, sambar, Muntjac and gaur. The line transect method was used to estimate absolute densities of the study species in November 2000. Distribution of the study species was studied using dung as an indicator of occupancy. Logistic regression models was used to examine species presence/absence and habitat parameters.Item Habitat Utilization by Hog Deer (Axis Porcinus) in Relation to other Sympatric Species at Jaldapara Wildlife Sanctuary(Wildlife Institute of India, Dehradun, 1999) Biswas, Tanushree; Mathur, V.B.; Sawarkar, V.B.This study on habitat utilization by hog deer (Axis porcinus) in relation to other sympatric species (chital & rhino) was carried out from December 1998 to April 1999 in Jaldapara Wildlife Sanctuary, West Bengal, The spatial and seasonal differences in habitat utilization of grasslands by hog deer, chital and rhino were investigated, The study was restricted to various types of grassland s existing within the sanctuary which were broadly stratified into five major types: Pure grassland (natural ), pure grassland (plantation), Savanna grassland (natural) and Savanna grassland ( plantation) and riverine grassland, Different habitat were traversed on riding elephants on selected paths to collect data on animal abundance/hr based on sightings in different habitat types, A total of nine transects were laid in different habitats to quantify availability of habitat parameters and their selection. Dung abundance was quantified using belt transects, at each 100m interval on the transects, The availability and utilization of habitats based on direct and Indirect methods were examined using Neu et al (1974), The results showed that hog deer selected areas with extensive grass cover with structural mosaics of cover and mostly early phenophase as opposed to habitats with low cover values, High abundance of hog deer were found in areas with higher proportion of Imperata cylindrica, Fire or burning of grasslands in combination with rain seemed to influence the overall selection of habitat by hog deer, Cutting of thatch by local people also influenced the habitat use by hog deer, Area preferred by hog deer were found to have high cattle dung density thus showing a very high pressure on areas preferred by hog deer, Interaction with chital seemed to be very low as the areas preferred by hogdeer are avoided by chital, Rhino also showed a similar trend with preference for plantation grassland areas and avoidance for natural grassland, Apart from the effect of these sympatric species directly, habitat use by hog deer seemed to have been influenced by the management steps taken to provide protection to Rhino, Plantation areas raised for the management of rhino were avoided by hog deer.