Theses and Dissertations

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    Habitat Utilisation of Asian Water Monitor (Varanus salvator) in and Around Bhitarkanika National Park, Odisha
    (Wildlife Institute of India, Dehradun, 2025) Rayen, W. Michael Isaac V.; Pandav, Bivash; Das, Abhijit
    The water monitor lizard Varanus salvator is the largest of all lizards in India, and its distribution is restricted to certain pockets of the northeast, eastern coast and Andaman and Nicobar Islands. Despite being the largest lizard, only limited studies have been done on this species, especially on the movement pattern and home range. This study focuses on the space use pattern of Varanus salvator in the mangrove habitats of Bhitarkanika National Park, Odisha. In total, five individuals were radio-tagged with VHF transmitters in the form of a backpack. All the tagged individuals were located twice in a day, and the location(coordinates), activity and other habitat variables were recorded. The data collected were analysed for their movement pattern, activity range and overlap between individuals and the human-dominated landscape was estimated using KDE. Along with the space use environmental variables (Ambient temperature, Relative humidity, vegetation species, canopy openness, etc) were analysed individually with behaviour to the relation between them using Fisher’s test. The results from this study add more information about its activity range, movement pattern and behavioural aspects in the mangrove landscape of India, which can be useful in the rescue, rehabilitation and management of the species.
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    Tigers in a multi-use forest: Prey, diet and conflict
    (Wildlife Institute of India, Dehradun, 2025) Chauhan, Aayush; Pandav, Bivash; Habib, Bilal
    This study provides a comprehensive assessment of tiger prey, dietary dynamics, and human-wildlife interactions in the Ramnagar Forest Division, a critical area outside protected areas in the Western Terai Arc. Despite a notable increase in tiger numbers—from 39 adults in 2015 to 67 in 2022—density surface modeling revealed persistently low populations of key wild prey species such as chital and sambar, primarily due to habitat degradation from historical timber-focused management. Diet analysis of genetically confirmed tiger scats demonstrated a strong reliance on large-bodied prey, with sambar and livestock comprising a significant portion of tiger biomass intake, reflecting both prey scarcity and ecological stress. The detection of plastic in both sambar and tiger digestive tracts further highlights the emerging threat of environmental pollution in multi-use forests. Human-tiger conflict remains acute, with 353 livestock depredation events and 28 attacks on people recorded over recent years, concentrated near settlements where prey and livestock overlap are highest. These findings underscore the urgent need for integrated conservation strategies that prioritize habitat restoration—such as grassland recovery—community-based conflict mitigation, improved waste management, and a shift in forest policy from timber production to biodiversity and coexistence. Only through such landscape-level, participatory approaches can the long-term viability of tigers and their prey be secured in shared, human-dominated areas like Ramnagar.
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    Ecology and ranging behaviour of Elephants, Elephas maximus, and its implications for managing human-elephant conflict in Chhattisgarh, India
    (Wildlife Institute of India, Dehradun, 2022) Lakshminarayanan, N.; Pandav, Bivash
    In the East-central region of India that supports the smallest regional elephant (Elephas maximus) population, the elephant ranges have expanded in northern Chhattisgarh (erstwhile eastern Madhya Pradesh) since late 1980s. Chhattisgarh was carved out of Madhya Pradesh during the year 2000, and thence, the State has been witnessing elephant range expansion and concomitant population growth resulting in acute human–elephant conflict. Over 60 human lives are lost every year due to human–elephant conflict, and the trend is only increasing. Chhattisgarh scenario represents challenges facing management of elephant populations undergoing environmental dispersals that typically result from saturated habitat conditions in the elephant home ranges. In response to the prevailing conflict situation, the study aimed to understand facets of elephant ecology and aspects of human–elephant conflict to generate management perspectives. Historic information available as fragments in literature indicates that elephant distribution in the whole of East-central region was marked by both range expansions and local extinctions during the last one century. In particular, the contemporary phenomenon of elephant range redistribution in the East-central region peaked after 1980s, and coincides with large-scale human-induced disturbances to the formerly intact elephant habitats in the region. There were also pull factors like creation of pseudo-habitats (providing only cover for elephants) through forestry operations that possibly attracted elephants to human–dominated areas perpetuating human-elephant conflict. In Chhattisgarh, the elephant distribution during the period 2012 to 2017 was reported from 16 Forest Divisions and four Protected Areas in the north and north-central regions of the state. The elephant population, as enumerated by Chhattisgarh Forest Department during 2021, ranged from 250 to 300.
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    Landscape level modeling of Asian Elephant (Elephas Maximus) habitat, Its population and interaction with humans in Nepal
    (Wildlife Institute of India, Dehradun, 2021) Ram, Ashok Kumar; Pandav, Bivash; Mondon, Samrat; Subedi, Naresh; Ram, Babu
    The nature and extent of human-elephant conflicts studied in Chitwan National Park and Parsa National Park complex (CNP & PNP) and its buffer zone in Nepal shows that crop damage and human casualties were more serious conflict issues among the other concerns of human–elephant conflict.
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    Determinants of occurrence of snow leopard and its prey species in the Indian Greater and Trans Himalaya.
    (Wildlife Institute of India, Dehradun, 2017) Ghoshal, Abhishek; Pandav, Bivash; Bhatanagar, Yash veer
    Understanding patterns of species occurrences and how their distribution and abundance are affected by abiotic and biotic factors have been a major theme of ecological research. The range and changes in the distribution of a species are important parameters to assess conservation status. Robust assessment of factors affecting species distribution and abundance contribute to the identification of conservation issues, appropriate scale(s) for species conservation and key stakeholders. This study is attempted to understand how distribution and abundance of a large-ranging and difficult-to-sample carnivore and its primary wild-prey species are affected by topographic, vegetation and land-use variables, at wide and fine spatial scales. Study is focused on the snow leopard (Panthera uncia), the elusive, rare and endangered apex predator of the Central and South Asian mountain systems, and its main prey, blue sheep or bharal (Pseudois nayaur) and Asiatic ibex (Capra sibirica) for the work. I have utilized tools from quantitative ecology and social sciences through the thesis.
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    Nesting ecology of colonial waterbirds at Bhitarkanika mangroves, Orissa
    (Wildlife Institute of India, Dehradun, 2010) Gopi, G.V.; Pandav, Bivash
    Colonial nesting is an important feature among a majority of the members of Pelicaniformes and Ciconiiformes. The nesting colonies of these birds that represent spatial and temporal clumping of nests are popularly referred to as heronries. One of the largest heronries in India is located in the Bhitarkanika mangroves, along the east coast of India. Birds use five species of mangrove trees for nesting which include Excoecaria agallocha (Guan), Heritiera fomes (Bada Sundari), Cynometra iripa (Singada), Hibiscus tiliaceus (Bania), Tamarix troupii (Jagula) for nesting in the heronry. . The breeding birds in this mixed species colony are Asian Openbill, Great Egret, Intermediate Egret, Little Egret, Cattle Egret, Grey Heron, Purple Heron, Black-crowned Night Heron, Little Cormorant, Darter, and Black-headed Ibis. The heronry in Bhitarkanika is located in an island covered with mangrove vegetation. The availability of large number of nest trees in terms of the mangrove forest and foraging areas in terms of the wetlands inside the forest and the paddy fields surrounding the forest are believed to be the factors favouring such large congregation of breeding water birds in the heronry
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    Occupancy and abundance of tigers and their prey in Terai Arc Landscape, Nepal.
    (Wildlife Institute of India, Dehradun, 2012) Karki, Jhamak Badur; Jhala, Y.V.; Pandav, Bivash
    The occupancy, distribution and tiger density are important information for management and conservation policy formulation. Realizing this need, this study was undertaken. The major wild prey species of tiger are spotted deer (Axis axis), wild pig (Sus scrofa), sambar (Rucervus unicolor), swamp deer (Rucervus duvaucelli duvaucelli), barking deer (Muntiacus muntjak), hog deer (Heylaphus porcinus) and gaur (Bos gaurus) in Nepal. To evaluate the occupancy of tiger in Nepal's Terai Arc landscape (TAL), sign survey was conducted in 96 grids (area 225 Km2 ) in 14 districts, including PAs. To assess correlation of the tiger occupancy with the availability of wild prey, and the human disturbance with the habitat use by the tiger, the sign of human disturbances (fire, timber cut, fuel wood collection, sign of poaching) and prey presence were recorded during the winter season. To estimate the density of tigers wild prey in PAs of Nepal's TAL, distance sampling was conducted along the predefined line transects during the summer season
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    Understanding aspects of demography and resource partitioning mechanisms of large felids among with abundance and spatial distribution of prey in Similipal Tiger Reserve, Odisha
    (Wildlife Institute of India, Dehradun, 2024) Rathore, Harshvardhan Singh; Pandav, Bivash; Jhala, Y.V.
    This study investigates the crucial aspect of coexistence for large predators like tigers and leopards sharing the same landscape and delves into the population dynamics of these animals by identifying the mechanisms of resource partitioning. By acting as a window into the inner workings of the ecosystem, the study reveals how tiger and leopard populations function alongside each other. Identifying the factors that influence their ability to share the resources successfully is vital for developing more effective conservation strategies. This knowledge ultimately contributes to ensuring the long-term survival of both tigers and leopards in this landscape. A precise assessment of several demographic parameters is necessary to make informed management decisions for their conservation within this unique habitat. These parameters include population density, sex structure, recruitment, and survival rates. With this background, the study comes with the following objectives. (1) To assess the abundance and spatial distribution of prey in STR.(2) To assess the various demographic parameters of large felids (tiger and leopard) in STR. (3) To determine the mechanisms of resource partitioning of the tiger and the leopard in STR. Similipal Tiger Reserve is located in the Mayurbhanj district of Odisha, between86˚06' and 86˚36' East longitude and 21˚31' and 22˚02' North latitude. The study was carried out over three years (2019–2021) across eight ranges in the core area of STR: Chahala, Nawana North, Nawana South, and Pithabata South Wildlife in the northern core, and UBK, Jenabil, Bhanjabasa, and National Park in the southern core.
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    Subsidized Mammals: Understanding the mammalian interactions with garbage sites around Western- Rajaji Tiger Reserve, Uttarakhand
    (Wildlife Institute of India, Dehradun, 2024) Nagarale, Shashank; Habib, Bilal; Shrotriya, Shivam; Pandav, Bivash
    Anthropogenic food subsidies in the form of garbage dumps are widespread across the globe. When such sites are found around protected areas/wildlife habitats, wildlife is observed to exploit these subsidies as food resources. Foraging on such human-derived foods may have a negative or positive impact on individual to community/ecosystem levels. It becomes important to know the species associated with garbage dumps when found around protected areas and understand the interactions happening at the garbage dumps. This study was conducted aiming to know the animals at risk and also increase our understanding of these novel setups. I deployed 13 IR camera traps at total 10 garbage sites resulting in a cumulative effort of 437 trap nights at the garbage site. The camera trap data was processed using Megadetector & SpSeg packages in python. This data was used to get the Relative Abundance Index with respect to 437 trap nights. Sambar (RAI= 8.35) was the species with the highest visitations at the garbage site, followed by Indian crested porcupine (RAI= 2.70) and spotted deer (RAI= 2.15). Synanthropes like rhesus macaque (RAI= 1.78 ) and wild pig (RAI= 1.67) were the most visitors following spotted deer. I hypothesised that synanthropic species will have higher visitations because of distribution outside protected areas, followed by omnivores from the forest area as compared to herbivores and carnivores from the forest areas because of their ability to exploit larger variety of food resources. The data from this study rejected this hypothesis as forest dwelling species like sambar and Indian crested porcupine visited these sites more as compared to other synanthropes. Lesser visitations by omnivores like golden jackal, small indian civet, small indian mongoose; carnivore like common leopard along with scavenger striped hyena was a result of the less percentage of poultry waste (0.33% at only one site) in the garbage sites. I collected data on variables such as distance of the garbage site from the protected area, human infrastructure and waterbody, also quantified the size of the garbage site in terms of perimeter, NDVI difference between the protected area and garbage site as a surrogate of vegetation cover, relative abundance of each species in the adjoining protected area and calculated the composite RAI values of human, dogs/cats and livestock as a disturbance measure . I quantified the garbage in broader categories using a 1 m2 quadrat which was divided into smaller sets of 25x25 cm2. A total of 39 plots were laid for all the garbage site. The categories quantified were food wrappers & food disposables, food materials, religious offerings, agriculture discards, dung plates, sanitary waste, construction waste, carcass & poultry waste, pharmaceutical waste, cloths and plastic and others. The relation between relative abundances within protected area and visitations at the garbage site was assessed using Spearman correlation test along with Wilcoxon test p-values. With the available dataset I used ANOSIM using ‘vegan’ package in R program v4.1.2 to investigate the effect by considering the dissimilarity of the communities between sites based on the RAI values of the species for each garbage site. Relative abundance from protected area was not related to visitation of the species at the garbage site. Out of the selected variables, the ANOSIM values suggested the effect of ‘distance between garbage site and protected area’ on forest species and synanthropes. The effect of the ‘size of garbage site’ on the species could not be tested as there was not much variation in the sizes of the garbage sites. Human, dogs/cats and livestock presence affected the forest species, suggesting their sensitivity towards disturbances, while synanthropes were found to have affinity with the NDVI difference between protected area and garbage sites (vegetation cover at the garbage site) and the ‘percentage of food materials available in the garbage site’. To understand the co-occurrences at the garbage site, I modified the probabilistic models for co-occurrences (Veech, 2013) to get pair-wise associations at the temporal scale of one hour. A total 13 mammals avoided co-occurring with humans (and vehicle) while nine with livestock and dogs. Co-occurrences were found to be there between sambar and Indian crested porcupine at two out of three religious sites and at a household site, suggesting high tolerance levels to each other’s presence at the garbage site to exploit the resources. One of the sites was visited by barking deer with spotted deer group/herd. This association by solitary living barking deer can be for foraging effectively by reducing vigilance efforts in a group.
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    Assessing the distribution and density of the Fishing Cat in Bhitarkanika mangroves of eastern India by
    (Wildlife Institute of India, Dehradun, 2024) Ashik, C.S.; Mukherjee, Shomita; Gopi, G.V.; Pandav, Bivash
    Identifying the areas of occurrence, population, and the factors affecting the species distribution is critical in conservation and management. The Fishing Cat (Prionailurus viverrinus), a smaller feline native to South and Southeast Asia, is globally vulnerable and adapted to wetland habitats, with a diet primarily consisting of fish. This makes Bhitarkanika, a mangrove ecosystem, an ideal habitat for the Fishing Cat. This study was conducted to estimate the species density, determine the factors affecting its distribution, and assess the effect of lunar illumination and tidal fluctuation on Fishing Cat activity. Despite known occurrences of Fishing Cats in this area, there has been no proper assessment of their population or other ecological studies. Camera traps were deployed in 109 grids, each covering 1km², across a total of 145 km², for 2878 trap nights. Camera trap locations are unbaited, with two cameras used to capture both flanks. Spatially explicit capture-recapture (SECR) was used to estimate the density, resulting in 0.6 ± 0.1 individuals per km². The abundance was also calculated, with an estimate of 99 ± 16 individuals. These density estimates from this study are close to recent high-density estimates for the species. The canopy cover, the width of the creek, canopy cover and the distance to the aquaculture farms significantly influenced the Fishing Cat habitat use, indicating the need for the conservation of natural mangrove habitats and regulation of aquafarms present near the forest. In contrast, the presence of Saltwater Crocodile showed a negative effect on the distribution of Fishing Cats. The Fishing Cats were more active during the brighter nights of the moon phase (μ= 266.783, r= 0.051, p< 0.01) due to better visual detection. The activity was influenced both by the tidal fluctuations in terms of food resources from waterbodies and it is further enhanced by the lunar illumination during the night. Pairwise non-parametric tests showed that the probability distributions of Fishing Cat activity under lunar illumination and tide fluctuation (D= 0.125, p-value = 0.9885 and D= 0.125, p-value= 0.9899) were similar, indicating minimal differences between each probability distribution. This study underscores the importance of protecting the mangrove habitat and controlling aquaculture expansion to ensure Fishing Cat survival in the Bhitarkanika National Park.