Theses and Dissertations

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    Tigers in a multi-use forest: Prey, diet and conflict
    (Wildlife Institute of India, Dehradun, 2025) Chauhan, Aayush; Pandav, Bivash; Habib, Bilal
    This study provides a comprehensive assessment of tiger prey, dietary dynamics, and human-wildlife interactions in the Ramnagar Forest Division, a critical area outside protected areas in the Western Terai Arc. Despite a notable increase in tiger numbers—from 39 adults in 2015 to 67 in 2022—density surface modeling revealed persistently low populations of key wild prey species such as chital and sambar, primarily due to habitat degradation from historical timber-focused management. Diet analysis of genetically confirmed tiger scats demonstrated a strong reliance on large-bodied prey, with sambar and livestock comprising a significant portion of tiger biomass intake, reflecting both prey scarcity and ecological stress. The detection of plastic in both sambar and tiger digestive tracts further highlights the emerging threat of environmental pollution in multi-use forests. Human-tiger conflict remains acute, with 353 livestock depredation events and 28 attacks on people recorded over recent years, concentrated near settlements where prey and livestock overlap are highest. These findings underscore the urgent need for integrated conservation strategies that prioritize habitat restoration—such as grassland recovery—community-based conflict mitigation, improved waste management, and a shift in forest policy from timber production to biodiversity and coexistence. Only through such landscape-level, participatory approaches can the long-term viability of tigers and their prey be secured in shared, human-dominated areas like Ramnagar.
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    Aspects and determinants of human-carnivore conflict in tiger corridors of Terai, Western Circle, Uttarakhand
    (Wildlife Institute of India, Dehradun, 2024) Ranjan, Vivek; Badola, Ruchi; Dhakate, Parag Madhukar
    Human-wildlife interaction with negative consequences, commonly termed Human- Wildlife Conflict (HWC), is a global conservation challenge. Understanding the cause-effect relationship resulting in HWC is essential to resolve and mitigate the conflict and promote human-wildlife coexistence. However, the increasing human population and rapidly changing demography severely affect the contiguity of forested landscapes and habitats, becoming a major challenge for conserving wide-ranging large mammals. The conservation and protection of wildlife corridors are vital for the survival of wide-ranging large mammals in the landscape. The Terai landscape, located in the foothills of the Himalayas and proximate plains of the Indian Himalayan Region, is one of the most significant and distinct transboundary landscapes worldwide. In recent decades, the Terai region has recorded a rapid increase in human population and industry growth due to its conducive topography for industrial development in the hilly state of Uttarakhand. The region is also home to diverse wildlife, and increased conservation efforts and protection have yielded positive results, with an increased population of endangered wildlife species like tigers in the last few years. With increasing population and shrinking wildlife habitat, human and wildlife's peaceful and sustainable coexistence is distressed, leading to an increased probability of negative human wildlife interaction. The study aims to assess the aspects and determinants of Human-Large Carnivore Conflict (HLCC) in the wildlife corridors and their adjoining habitats in the eastern terai landscape of Uttarakhand state of India. The large carnivores considered for the purpose of this study are tiger and leopard only, which are common in the landscape of the study area. The study has three objectives: to assess the nature and extent of human-wildlife conflict due to large carnivores, the habitat structure and composition of the wildlife corridors and the diet preferences and consumption patterns of large carnivores in the corridor habitat. The thesis has been divided into six chapters, where the first two chapters are a general introduction and literature review, and the following four chapters talk about socio-ecological factors, habitat quality of corridor habitats, implications of habitat changes on HLCC, and diet profile of two sympatric large carnivores of the study area, i.e. tiger and leopard. The study area is part of Terai Arc Landscape (TAL), a Tiger Conservation Landscape (TCL_Id-44) of global importance with the potential for increasing the wild tiger population. The study was conducted in the delineated wildlife corridors for large mammals in the terai landscape of Uttarakhand state of India. The study focuses on the critical wildlife corridor in high tiger density and transboundary areas, providing crucial connectivity of habitats and wildlife populations. The study area is divided into two study blocks for the spatial convenience of the study: Block 1 comprises the Kosi Corridor, and Block 2 comprises theKilpura-Khatima-Surai (KKS) and Boom-Brahmadey (BB) corridor and their adjoining forests. Kosi corridor connects Corbett Tiger Reserve to Pawalgarh Conservation Reserve in Ramnagar forest division. The KKS and BB corridors in Block 2 have contiguous and interconnected habitats adjoining Nandhaur Wildlife Sanctuary in the Haldwani Forest Division of Uttarakhand, connecting to Pilibhit Tiger Reserve and Sukhlaphanta National Park through forest of Nepal. The study area is rich in faunal diversity and home to a few important large mammals, such as the Tiger, Leopard and the Asian elephant. In order to understand the nature and extent of HLCC in the landscape, it was essential to understand the socio-economic conditions and perceptions of the communities. We conducted random household (HH) semi-structured questionnaire surveys and discussions in the villages located in the corridors. The information on HWC compensation records was also collected from the Uttarakhand Forest department to analyze the patterns of conflict incidences. The results show that livestock grazing, fuelwood, and fodder collection from the forest are common and prevalent practices. About 90% of the households have LPG connections at home, yet most HH members still visit forests for collection. The education level among the respondents is low, with maximum HHs having an annual income of less than one lakh. The major occupation as the primary source of income in the study area is casual labour work, and most of the HHs have land holdings of less than an acre. The incidences of HLCC mainly occurred inside the forest areas and later half of the daytime, i.e., from noon to evening. The season of incidence and condition of the body of the depredated livestock show significant association with the place of incidence. The seasonality pattern of HLCC incidences shows nan incidents during monsoon in study block 1, whereas in block 2, it is in winter. The trend of incidences over the last 13 years in study block 1 shows a peak in tiger and leopard cases during July and August and forecasts an increase in cases over upcoming years. The annual trend of HLCC in the two study blocks is significantly different. The nature of HLCC in the study area is both tangible, which can be visually observed and intangible, which has imbibed effects with direct and indirect impacts. The tangible nature of large carnivore conflict is the direct impacts in the form of human casualty and livestock depredation. The intangible nature of the conflict with indirect impacts are psychological— fear of large carnivores and sense of insecurity, social — loss of livelihood and family security. The aspects of HLCC are socio-ecological, with both the social factors of communities residing around the forest and the ecological processes influencing the HWI. In rapidly transforming human-dominated landscapes, anthropogenic activities and disturbances significantly impact the ecological processes and landscape characteristics. The habitat quality was assessed using the nested quadrat method for vegetation and camera trapping for large carnivore population estimation and prey availability. The camera trapping was done using the capture-recapture method in 2 sq. Km sampling grids. The results show maximum tree diversity in the BB corridor, with good to fair Sal regeneration in all three corridors. However, most of the tree species show poor to no regeneration. Sixty-one species of trees were recorded from the three corridor areas sampled. As an invasive plant species, Lantana camara is more prevalent and extensive in study block 1 and Ageratina adenophara in block 2. The grassland habitat is shrinking, and less in all three corridor habitats.Anthropogenic disturbance is high in block 2 in comparison to block 1, while tree cutting and lopping are the most prevalent anthropogenic disturbance signs observed in all three corridors at equivalent levels. The medium-sized prey is the most abundant prey base in both the study blocks, with maximum relative abundance of Spotted deer. However, the relative abundance of Sambar deer is higher in the bhabar topography area of corridors and maximum in large sized prey. The study block | has 16 adult tigers and 22 leopards, while block 2 has 31 adult tigers and 40 leopards. The wildlife habitats of the three corridor areas studied have good tree diversity and complexity. However, the regeneration of tree species is poor for most species. The shrub and herb cover are also low for most areas in all three corridors, affecting the lower structure of forest habitats. The extensive growth of weed species like Lantana camara and Ageratina adenophora also affects understory species' composition, structure, and tree regeneration. The poor regeneration and deteriorated lower strata vegetation with sparse undergrowth are unfavourable conditions for breeding large carnivores, which requires dense patches for hiding their offspring. The lack of such patches and high human disturbance forces them out of the forest to use sugarcane farm fields to raise and hide their young ones, as has been observed in many instances in the Terai-Bhabar landscape. Using human-modified landscapes by large carnivores for their life cycle stages is unsuitable for human-wildlife coexistence and increases the vulnerability of negative HWI. ASS Bind Expert in CC EE TTT The implications of habitat transformations on HLCC were assessed by identifying spatial conflict hotspots based on compensation records of the last 14 years and the magnitude of incidents related to large carnivores in different locations. The Land Use Land Cover (LULC) and Enhanced Vegetation Index (EVI) change over twenty years were assessed. The Nighttime Light (NTL), an indicator of human growth and activity centres, was also assessed for a decadal change. The impact of changes in these habitat attributes on hotspots was assessed using the Generalized Linear Model. The model shows that the impact of vegetation health change and human activity or disturbances have a significant effect on conflict hotspots. The very high-risk zones are also located in the delineated corridors near bottleneck areas. The study has highlighted that the implications of habitat changes are significant for HLCC. The implications of habitat changes vary with the landscape and regional attributes, as observed in our study at two sites with varied underlying factors other than habitats. In the present conservation paradigm of large carnivores, it is essential to understand the feeding habits and prey-predator dynamics outside protected areas. Our study fills this gap in our understanding of the dietary habits of sympatric large carnivore species outside protected areas (PAs) in the terai landscape of India. It focuses on the feeding ecology of tigers and leopards in wildlife corridor habitats outside PAs. Our results show a significant dependence of tigers and leopards on medium-sized prey, i.e., Wild boars and spotted deer. It also shows a 93% overlap between tiger and leopard diets in the study area. However, it is crucial to augment wild prey availability in the wildlife corridor habitats, where grazing pressure is high. The problem of stray cattle has further escalated the issue of livestock depredation by large carnivores. Conservation strategists must consider the feeding habits of the predators and how these are changing due to the induced effects of anthropogenic activities.
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    Tiger and leopard: Population ecology and resource portioning of sympatric carnivores in Kanha Tiger Reserve. M.P.
    (wildlife Institute of India, Dehradun, 2020) Kumar, Ujjwal; Jhala, Y.V.; Qureshi, Qamar
    Large investment in the form of protected areas, human resettlement, law enforcement-patrols and habitat management are made to conserve tigers and reap their benevolent umbrella role to secure India’s biodiversity. The objectives of present study are (i) the trend of tiger and leopard population (ii) demography of tigers and leopards and mechanism of their co existence. The study area was Kanha tiger reserve (area 2074 km2), a major stronghold for both species in central Indian landscape. Study investigate the estimated vital parameters of tiger demography through robust design based classical open population models in which population was open to demographic process. This study used both camera trap based mark-recapture and continuous monitoring known fate information to estimate reproductive parameters along with age specific survival and dispersal probability of tigers.
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    Movement and space use pattern of Tigers in the Eastern Vidarbha landscape, Maharashtra, India
    (Wildlife Institute of India, Dehradun, 2024) Hussain, Zehidul; Habib, Bilal
    Movement is a crucial ecological process that influences the survival, population dynamics, and evolution of animals. Animal movement patterns are diverse and are influenced by life stages, sex, age, and habitat requirements. The relationship between animal movement and conservation is complex, particularly due to human-induced environmental changes This study emphasises that various factors influence animal movement, including individual characteristics and external factors such as resource availability, distribution, competition, and predation. Animals adjust their space use based on seasonal changes in resource availability. Additionally, movement patterns and space use can be influenced by habitat disruption, forcing individuals to displace and establish new home ranges. Landscape features play a significant role in animal home ranges, with resource distribution determining the extent of the required area. Fragmentation of habitats and human activities can affect landscape connectivity, leading to isolation, reduced gene flow, and potential genetic bottlenecks or local extinctions. Thus, maintaining habitat connectivity is crucial for long-term survival of highly mobile species. The study highlights the need for comprehensive datasets and technological advancements, such as GPS tracking, to study animal movement and behaviour. It also emphasises the importance of studying dispersal patterns, as dispersed individuals often encounter humandominated landscapes and face increased vulnerability to anthropogenic effects. This study focused on quantifying movement patterns and space use across three life stages: predispersal, dispersal, and post-dispersal phases. The results revealed differences in space use within and outside protected areas, with individuals outside protected areas requiring larger territories. The study also identified seasonal variations in space use and movement patterns, with dispersed tigers exhibiting the highest space use. Land-use composition, landscape configuration, and anthropogenic disturbances influence the use of tiger space. These findings provide insights into the factors influencing tiger movement and space use in a human-dominated landscape. They highlighted the importance of maintaining habitat connectivity, reducing anthropogenic disturbances, and restoring suitable habitats outside protected areas to facilitate dispersal and ensure the long-term survival of tigers.
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    Meta-population Dynamics of Tiger (Panthera Tigris Tigris) in the Terai-Arc Landscape, India
    (Wildlife Institute of India, Dehradun, 2021) Biswas, Suvankar; Mondol, Samrat
    This thesis focused to add deeper understanding of populations at a large, contiguous landscape Studying the tiger source-sink population dynamics in relation to current habitat scenarios would be crucial to set the site-specific future management strategies for successful tiger conservation mitigation. In this thesis, a combination of intensive field surveys, non-invasive genetic tools and GIS modelling were used to assess the tiger population connectivity across the TAL. More specifically, genetic diversity, population structure, demographic changes, source and sink populations and the habitat features which help to retain such population dynamics were investigated. Objectives: This thesis has three objecives which have formed three separate chapters. i. Population structure and demographic history of tigers in the Terai-Arc landscape, India. ii. Source-sink dynamics of tigers across different protected and nonprotected habitats in the Terai-Arc landscape, India. iii. Assessment of landscape connectivity of tigers in the Terai-Arc landscape, India.
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    Understanding aspects of demography and resource partitioning mechanisms of large felids among with abundance and spatial distribution of prey in Similipal Tiger Reserve, Odisha
    (Wildlife Institute of India, Dehradun, 2024) Rathore, Harshvardhan Singh; Pandav, Bivash; Jhala, Y.V.
    This study investigates the crucial aspect of coexistence for large predators like tigers and leopards sharing the same landscape and delves into the population dynamics of these animals by identifying the mechanisms of resource partitioning. By acting as a window into the inner workings of the ecosystem, the study reveals how tiger and leopard populations function alongside each other. Identifying the factors that influence their ability to share the resources successfully is vital for developing more effective conservation strategies. This knowledge ultimately contributes to ensuring the long-term survival of both tigers and leopards in this landscape. A precise assessment of several demographic parameters is necessary to make informed management decisions for their conservation within this unique habitat. These parameters include population density, sex structure, recruitment, and survival rates. With this background, the study comes with the following objectives. (1) To assess the abundance and spatial distribution of prey in STR.(2) To assess the various demographic parameters of large felids (tiger and leopard) in STR. (3) To determine the mechanisms of resource partitioning of the tiger and the leopard in STR. Similipal Tiger Reserve is located in the Mayurbhanj district of Odisha, between86˚06' and 86˚36' East longitude and 21˚31' and 22˚02' North latitude. The study was carried out over three years (2019–2021) across eight ranges in the core area of STR: Chahala, Nawana North, Nawana South, and Pithabata South Wildlife in the northern core, and UBK, Jenabil, Bhanjabasa, and National Park in the southern core.
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    Spatial and Temporal Responses of Leopard (Panthera pardus) in the Presence of Tiger (Panthera tigris tigris) in Rajaji National Park Uttarakhand
    (2021) Krishna, Anubhuti; Lyngdoh, Salvador
    Leopards co-occur with tigers in several parts of Asia as well as over a major portion of their geographical distribution in India. A clear dominance hierarchy establishes between the two felid species in regions of sympatry with tigers, owing to stark body size differences, holding higher ground. The striped felid, thus, invokes numerical as well as functional responses in their less dominant counterparts. The current study aims at understanding the spatial and temporal patterns of leopards in the presence of tigers. Rajaji National Park in Uttarakhand offers an excellent natural setup for understanding such intra-guild interactions as the western part of the Park hosts leopards but is devoid of tigers and the eastern region has sympatric populations of the two felid species. These act as control and experimental setups respectively, and hence this site was chosen for conducting the field sampling. It was found that leopards exhibit some degree of spatial segregation from tigers as was evidenced by differences in the activity hotspots of the two felids in Eastern Rajaji. This is attributable, at least in part, to tigers as a negative relationship was obtained between space use by tigers and leopard occupancy under the occupancy framework. Temporal patterns, on the other hand, revealed a rather interesting trend. The diel activity of leopards in Eastern Rajaji, i.e., in the presence of tigers was very different from that of leopards in the western part of the Park. The activity of the latter was distributed throughout the day with nocturnal peaks of low intensity. The former, however, showed an activity pattern restricted to the nocturnal hours with very little activity during the day. The temporal peaks for this population also showed greater intensity than that of the pattern observed in the western leopard population.These peaks of activity, however, contrary to expectations, coincided with those of tigers in the landscape. Since predator-prey interactions also play an important role in shaping the temporal activity patterns of carnivores, the temporal overlaps between the two felids and their potential prey was also analysed. This too failed to offer an explanation to the rather unexpected leopard diel pattern in ER. It was subsequently observed that though leopards might have similar temporal activity to that of tigers, they are rarely at the same place at the same time. A clear lag was observed between leopard captures at camera traps following photo-captures of tigers at the same location. Hence, a combination of spatial and temporal segregation might be at play at a finer scale than at the population level and requires further examination in order to elucidate the interactions and mechanisms of sympatry between leopards and tigers in this landscape
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    Playing it Safe Tiger Movement and Livestock Depredation Patterns in Panna Tiger Reserve, Central India
    (Wildlife Institute of India, Dehradun, 2021) Mohan, Manu K.; Ramesh, K.; Sathyakumar, S.
    The movement of large carnivores in human-dominated landscapes is not uncommon anymore. While their presence is superficially tolerated or ignored, it is a consistent cause of concern to the people living in its proximity, due to competing interests in sharing of resources and direct threat to human lives. Carnivores incur energetic costs in the form of restricted activity and having to make spatial and temporal activity adjustments. Humans, on the other hand, face losses from livestock depredation and/or loss of lives. Depredation patterns of livestock by carnivores are associated with a variety of factors including habitat structure as well as movement patterns of carnivores in relation to anthropogenic and livestock activity. Hence, deducing movement patterns of the groups in conflict is an important step in understanding the dynamics of conflict patterns at fine scales. Supplemented by models to predict the probability of conflicts spatially which delineates conflict hotspots, site-specific mitigation measures could be devised. To understand such patterns of carnivore movement and relation to livestock depredation events, I conducted this study in a human-dominated region adjacent to the Critical Tiger Habitat of Panna Tiger Reserve in central India from December 2020 to April 2021, within an area of 200 km2. The broad objective was to model the conflict probability based on the proportion of the human-dominated area used by tigers, its intensity of use and spatio-temporal activity within the area, in response to human disturbances. For this, I used intensive camera trap sampling at fine-scale within grids (1 km x 1 km) across the entire study area. Movement parameters of tigers such as displacement and distance to villages in day and night were also analysed by monitoring two GPS-collared tigers in the study area. Occupancy estimates showed 74% of the sampled area being used by tigers, with the relative abundance of a large bodied wild prey (sambar) as the strongest predictor. In addition, a clear positive trend in the relative abundance index of tigers with distance to villages was observed. The temporal activities of the tigers also showed a significant difference in median activity time, shifting towards the night along with moderate diurnal activity at higher distances from villages. The tigers exhibited both crepuscular and nocturnal activity at close distances to villages. The movement range analyses of the tigers using camera traps showed the study area being used by 17 tigers, with 9 of them being adults (>3 years of age). Fixed Kernel Density home-range estimates of the two GPS-collared tigers showed a considerable proportion of their home range areas falling within village boundaries (10%) and an average of 16% of all GPS locations of the tigers fell within 0-100 metres from villages. Despite such high exploratory use of areas within and close distances to villages by sub-adults, there were relatively low incidences of conflict between the tigers and villagers in the study area. Finally, to assess the extent of risk and predict the probability of livestock depredation in the area, Generalized Linear Models were used. The best-fit model indicated a significant negative effect of distance to villages and highway to the probability of livestock depredation. Linear intrusions through highway and forest trails from villages allow easy access to livestock and herders for the utilization of the multi-use buffer areas of the Reserve as well as the tigers to come in proximity to settlements. Corralling of livestock in the night, avoiding free-ranging and grazing of livestock in the identified conflict hotspots, incentivization of alternate source of livelihood by strengthening the institutions such as Eco-Development Committees, along with continuous monitoring of tigers which range close to human settlements are discussed as some of the important measures which can be adopted to decrease the interfaces leading to tiger-human conflict in Panna Tiger Reserve.