Theses and Dissertations
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Item Spatial pattern of termite mounds and its association with vegetation with experimental insights on mound repair in response to ant diversity(Wildlife Institute of India, Dehradun, 2025) Vishvavikash, I.K.Termites are considered pests all around the world in Human settlements and agricultural fields. But in Nature, termites are ecosystem engineers, they play a key role in keeping the ecosystem balanced by processing plant material and returning nutrients to the environment. In the African savanna and the Tropical Rainforest of Malaysia, termites are well studied, and termite mounds are considered nutritional hotspots. In the African continent, the genus Macrotermes is well studied, and their distribution mainly depends on the colony size. The larger colony size utilizes more space for foraging around the mound and also shows great intra- and interspecific competition between the colonies. In India, very few studies have been conducted on Termites. In this study, we investigated the spatial ecology of Termite mounds (genus Odontotermes) in the Sal-dominant Forest of Western Rajaji National Park. To understand the spatial ecology of termite mounds, we assessed their spatial distribution patterns using GPS mapping, mound morphometric characteristics, the influence of surrounding vegetation, and soil chemical properties. We studied the mound repair dynamics of the termites with respect to the predatory pressure exerted by the surrounding mound organisms. We created artificial breaches in termite mounds to monitor repair rates, while pitfall traps around mounds assessed ant diversity and abundance in the surrounding area. The Size classes of the Termite Mound didn’t show any kind of territoriality around them. Vegetation around termite mounds (Odontotermes) didn’t show any type of heterogeneity around the termite mounds. Termite mounds of the small size class show a faster repair rate than the larger termite mounds. The Ant diversity does not dictate the termite mound repair rate.Item Habitat Characterization and Spatial Distribution of Quercus lanata Sm. in the Kumaon Himalaya(Wildlife Institute of India, Dehradun, 2025) Kaushik, Shikhar; Kumar, Amit; Rawat, G.S.Oaks are climax species which evolved around 56 million years ago. India is home to around 35 species of oaks out of which only 5 species are native to Uttarakhand. In Western Himalaya 3 species make extensive continuous forests which includes Banj oak forest, Moru oak forest and Kharsu oak forest. They show gregarious nature and provide countless ecological and economic benefits to the country. The other two species, Quercus glauca and Quercus lanata are patchy is distribution. For Quercus lanata the Kumaon Himalaya is the western most limit and therefore that species is patchy and shows disjunct population. This study aimed to understand the population, regeneration, habitat characterisation and distribution of Quercus lanata in Kumaon. Descriptive analysis was used to understand the population structure and associate species and maxent modelling was used to develop habitat suitability model. The overall population of Quercus lanata (Rianj) in Kumaon shows a stable and gradually growing trend. Quercus lanata shows a diverse preference to habitat which ranges from northerns moist slopes to drier southern slopes. The significant values from analysed data suggest that its preference at southern slope is comparatively higher. It occurs with Quercus leucotrichophora as well as with Pinus roxburghii. The major co-occurring species with Quercus lanata are Rhododendron arboreum and Myrica esculenta. The developed habitat suitability and generalised linear models helped to understand that elevation and aspect play a significant role in distribution of Quercus lanata. The demand of Quercus lanata for fodder and fuelwood in village area is quite high and therefore requires sustainable practices. As of now Quercus lanata is classified under either Banj oak forest or Moru oak forest which reduces the level of conservation and management this species require.Item Tigers in a multi-use forest: Prey, diet and conflict(Wildlife Institute of India, Dehradun, 2025) Chauhan, Aayush; Pandav, Bivash; Habib, BilalThis study provides a comprehensive assessment of tiger prey, dietary dynamics, and human-wildlife interactions in the Ramnagar Forest Division, a critical area outside protected areas in the Western Terai Arc. Despite a notable increase in tiger numbers—from 39 adults in 2015 to 67 in 2022—density surface modeling revealed persistently low populations of key wild prey species such as chital and sambar, primarily due to habitat degradation from historical timber-focused management. Diet analysis of genetically confirmed tiger scats demonstrated a strong reliance on large-bodied prey, with sambar and livestock comprising a significant portion of tiger biomass intake, reflecting both prey scarcity and ecological stress. The detection of plastic in both sambar and tiger digestive tracts further highlights the emerging threat of environmental pollution in multi-use forests. Human-tiger conflict remains acute, with 353 livestock depredation events and 28 attacks on people recorded over recent years, concentrated near settlements where prey and livestock overlap are highest. These findings underscore the urgent need for integrated conservation strategies that prioritize habitat restoration—such as grassland recovery—community-based conflict mitigation, improved waste management, and a shift in forest policy from timber production to biodiversity and coexistence. Only through such landscape-level, participatory approaches can the long-term viability of tigers and their prey be secured in shared, human-dominated areas like Ramnagar.Item Aspects and determinants of human-carnivore conflict in tiger corridors of Terai, Western Circle, Uttarakhand(Wildlife Institute of India, Dehradun, 2024) Ranjan, Vivek; Badola, Ruchi; Dhakate, Parag MadhukarHuman-wildlife interaction with negative consequences, commonly termed Human- Wildlife Conflict (HWC), is a global conservation challenge. Understanding the cause-effect relationship resulting in HWC is essential to resolve and mitigate the conflict and promote human-wildlife coexistence. However, the increasing human population and rapidly changing demography severely affect the contiguity of forested landscapes and habitats, becoming a major challenge for conserving wide-ranging large mammals. The conservation and protection of wildlife corridors are vital for the survival of wide-ranging large mammals in the landscape. The Terai landscape, located in the foothills of the Himalayas and proximate plains of the Indian Himalayan Region, is one of the most significant and distinct transboundary landscapes worldwide. In recent decades, the Terai region has recorded a rapid increase in human population and industry growth due to its conducive topography for industrial development in the hilly state of Uttarakhand. The region is also home to diverse wildlife, and increased conservation efforts and protection have yielded positive results, with an increased population of endangered wildlife species like tigers in the last few years. With increasing population and shrinking wildlife habitat, human and wildlife's peaceful and sustainable coexistence is distressed, leading to an increased probability of negative human wildlife interaction. The study aims to assess the aspects and determinants of Human-Large Carnivore Conflict (HLCC) in the wildlife corridors and their adjoining habitats in the eastern terai landscape of Uttarakhand state of India. The large carnivores considered for the purpose of this study are tiger and leopard only, which are common in the landscape of the study area. The study has three objectives: to assess the nature and extent of human-wildlife conflict due to large carnivores, the habitat structure and composition of the wildlife corridors and the diet preferences and consumption patterns of large carnivores in the corridor habitat. The thesis has been divided into six chapters, where the first two chapters are a general introduction and literature review, and the following four chapters talk about socio-ecological factors, habitat quality of corridor habitats, implications of habitat changes on HLCC, and diet profile of two sympatric large carnivores of the study area, i.e. tiger and leopard. The study area is part of Terai Arc Landscape (TAL), a Tiger Conservation Landscape (TCL_Id-44) of global importance with the potential for increasing the wild tiger population. The study was conducted in the delineated wildlife corridors for large mammals in the terai landscape of Uttarakhand state of India. The study focuses on the critical wildlife corridor in high tiger density and transboundary areas, providing crucial connectivity of habitats and wildlife populations. The study area is divided into two study blocks for the spatial convenience of the study: Block 1 comprises the Kosi Corridor, and Block 2 comprises theKilpura-Khatima-Surai (KKS) and Boom-Brahmadey (BB) corridor and their adjoining forests. Kosi corridor connects Corbett Tiger Reserve to Pawalgarh Conservation Reserve in Ramnagar forest division. The KKS and BB corridors in Block 2 have contiguous and interconnected habitats adjoining Nandhaur Wildlife Sanctuary in the Haldwani Forest Division of Uttarakhand, connecting to Pilibhit Tiger Reserve and Sukhlaphanta National Park through forest of Nepal. The study area is rich in faunal diversity and home to a few important large mammals, such as the Tiger, Leopard and the Asian elephant. In order to understand the nature and extent of HLCC in the landscape, it was essential to understand the socio-economic conditions and perceptions of the communities. We conducted random household (HH) semi-structured questionnaire surveys and discussions in the villages located in the corridors. The information on HWC compensation records was also collected from the Uttarakhand Forest department to analyze the patterns of conflict incidences. The results show that livestock grazing, fuelwood, and fodder collection from the forest are common and prevalent practices. About 90% of the households have LPG connections at home, yet most HH members still visit forests for collection. The education level among the respondents is low, with maximum HHs having an annual income of less than one lakh. The major occupation as the primary source of income in the study area is casual labour work, and most of the HHs have land holdings of less than an acre. The incidences of HLCC mainly occurred inside the forest areas and later half of the daytime, i.e., from noon to evening. The season of incidence and condition of the body of the depredated livestock show significant association with the place of incidence. The seasonality pattern of HLCC incidences shows nan incidents during monsoon in study block 1, whereas in block 2, it is in winter. The trend of incidences over the last 13 years in study block 1 shows a peak in tiger and leopard cases during July and August and forecasts an increase in cases over upcoming years. The annual trend of HLCC in the two study blocks is significantly different. The nature of HLCC in the study area is both tangible, which can be visually observed and intangible, which has imbibed effects with direct and indirect impacts. The tangible nature of large carnivore conflict is the direct impacts in the form of human casualty and livestock depredation. The intangible nature of the conflict with indirect impacts are psychological— fear of large carnivores and sense of insecurity, social — loss of livelihood and family security. The aspects of HLCC are socio-ecological, with both the social factors of communities residing around the forest and the ecological processes influencing the HWI. In rapidly transforming human-dominated landscapes, anthropogenic activities and disturbances significantly impact the ecological processes and landscape characteristics. The habitat quality was assessed using the nested quadrat method for vegetation and camera trapping for large carnivore population estimation and prey availability. The camera trapping was done using the capture-recapture method in 2 sq. Km sampling grids. The results show maximum tree diversity in the BB corridor, with good to fair Sal regeneration in all three corridors. However, most of the tree species show poor to no regeneration. Sixty-one species of trees were recorded from the three corridor areas sampled. As an invasive plant species, Lantana camara is more prevalent and extensive in study block 1 and Ageratina adenophara in block 2. The grassland habitat is shrinking, and less in all three corridor habitats.Anthropogenic disturbance is high in block 2 in comparison to block 1, while tree cutting and lopping are the most prevalent anthropogenic disturbance signs observed in all three corridors at equivalent levels. The medium-sized prey is the most abundant prey base in both the study blocks, with maximum relative abundance of Spotted deer. However, the relative abundance of Sambar deer is higher in the bhabar topography area of corridors and maximum in large sized prey. The study block | has 16 adult tigers and 22 leopards, while block 2 has 31 adult tigers and 40 leopards. The wildlife habitats of the three corridor areas studied have good tree diversity and complexity. However, the regeneration of tree species is poor for most species. The shrub and herb cover are also low for most areas in all three corridors, affecting the lower structure of forest habitats. The extensive growth of weed species like Lantana camara and Ageratina adenophora also affects understory species' composition, structure, and tree regeneration. The poor regeneration and deteriorated lower strata vegetation with sparse undergrowth are unfavourable conditions for breeding large carnivores, which requires dense patches for hiding their offspring. The lack of such patches and high human disturbance forces them out of the forest to use sugarcane farm fields to raise and hide their young ones, as has been observed in many instances in the Terai-Bhabar landscape. Using human-modified landscapes by large carnivores for their life cycle stages is unsuitable for human-wildlife coexistence and increases the vulnerability of negative HWI. ASS Bind Expert in CC EE TTT The implications of habitat transformations on HLCC were assessed by identifying spatial conflict hotspots based on compensation records of the last 14 years and the magnitude of incidents related to large carnivores in different locations. The Land Use Land Cover (LULC) and Enhanced Vegetation Index (EVI) change over twenty years were assessed. The Nighttime Light (NTL), an indicator of human growth and activity centres, was also assessed for a decadal change. The impact of changes in these habitat attributes on hotspots was assessed using the Generalized Linear Model. The model shows that the impact of vegetation health change and human activity or disturbances have a significant effect on conflict hotspots. The very high-risk zones are also located in the delineated corridors near bottleneck areas. The study has highlighted that the implications of habitat changes are significant for HLCC. The implications of habitat changes vary with the landscape and regional attributes, as observed in our study at two sites with varied underlying factors other than habitats. In the present conservation paradigm of large carnivores, it is essential to understand the feeding habits and prey-predator dynamics outside protected areas. Our study fills this gap in our understanding of the dietary habits of sympatric large carnivore species outside protected areas (PAs) in the terai landscape of India. It focuses on the feeding ecology of tigers and leopards in wildlife corridor habitats outside PAs. Our results show a significant dependence of tigers and leopards on medium-sized prey, i.e., Wild boars and spotted deer. It also shows a 93% overlap between tiger and leopard diets in the study area. However, it is crucial to augment wild prey availability in the wildlife corridor habitats, where grazing pressure is high. The problem of stray cattle has further escalated the issue of livestock depredation by large carnivores. Conservation strategists must consider the feeding habits of the predators and how these are changing due to the induced effects of anthropogenic activities.Item Subsidized Mammals: Understanding the mammalian interactions with garbage sites around Western- Rajaji Tiger Reserve, Uttarakhand(Wildlife Institute of India, Dehradun, 2024) Nagarale, Shashank; Habib, Bilal; Shrotriya, Shivam; Pandav, BivashAnthropogenic food subsidies in the form of garbage dumps are widespread across the globe. When such sites are found around protected areas/wildlife habitats, wildlife is observed to exploit these subsidies as food resources. Foraging on such human-derived foods may have a negative or positive impact on individual to community/ecosystem levels. It becomes important to know the species associated with garbage dumps when found around protected areas and understand the interactions happening at the garbage dumps. This study was conducted aiming to know the animals at risk and also increase our understanding of these novel setups. I deployed 13 IR camera traps at total 10 garbage sites resulting in a cumulative effort of 437 trap nights at the garbage site. The camera trap data was processed using Megadetector & SpSeg packages in python. This data was used to get the Relative Abundance Index with respect to 437 trap nights. Sambar (RAI= 8.35) was the species with the highest visitations at the garbage site, followed by Indian crested porcupine (RAI= 2.70) and spotted deer (RAI= 2.15). Synanthropes like rhesus macaque (RAI= 1.78 ) and wild pig (RAI= 1.67) were the most visitors following spotted deer. I hypothesised that synanthropic species will have higher visitations because of distribution outside protected areas, followed by omnivores from the forest area as compared to herbivores and carnivores from the forest areas because of their ability to exploit larger variety of food resources. The data from this study rejected this hypothesis as forest dwelling species like sambar and Indian crested porcupine visited these sites more as compared to other synanthropes. Lesser visitations by omnivores like golden jackal, small indian civet, small indian mongoose; carnivore like common leopard along with scavenger striped hyena was a result of the less percentage of poultry waste (0.33% at only one site) in the garbage sites. I collected data on variables such as distance of the garbage site from the protected area, human infrastructure and waterbody, also quantified the size of the garbage site in terms of perimeter, NDVI difference between the protected area and garbage site as a surrogate of vegetation cover, relative abundance of each species in the adjoining protected area and calculated the composite RAI values of human, dogs/cats and livestock as a disturbance measure . I quantified the garbage in broader categories using a 1 m2 quadrat which was divided into smaller sets of 25x25 cm2. A total of 39 plots were laid for all the garbage site. The categories quantified were food wrappers & food disposables, food materials, religious offerings, agriculture discards, dung plates, sanitary waste, construction waste, carcass & poultry waste, pharmaceutical waste, cloths and plastic and others. The relation between relative abundances within protected area and visitations at the garbage site was assessed using Spearman correlation test along with Wilcoxon test p-values. With the available dataset I used ANOSIM using ‘vegan’ package in R program v4.1.2 to investigate the effect by considering the dissimilarity of the communities between sites based on the RAI values of the species for each garbage site. Relative abundance from protected area was not related to visitation of the species at the garbage site. Out of the selected variables, the ANOSIM values suggested the effect of ‘distance between garbage site and protected area’ on forest species and synanthropes. The effect of the ‘size of garbage site’ on the species could not be tested as there was not much variation in the sizes of the garbage sites. Human, dogs/cats and livestock presence affected the forest species, suggesting their sensitivity towards disturbances, while synanthropes were found to have affinity with the NDVI difference between protected area and garbage sites (vegetation cover at the garbage site) and the ‘percentage of food materials available in the garbage site’. To understand the co-occurrences at the garbage site, I modified the probabilistic models for co-occurrences (Veech, 2013) to get pair-wise associations at the temporal scale of one hour. A total 13 mammals avoided co-occurring with humans (and vehicle) while nine with livestock and dogs. Co-occurrences were found to be there between sambar and Indian crested porcupine at two out of three religious sites and at a household site, suggesting high tolerance levels to each other’s presence at the garbage site to exploit the resources. One of the sites was visited by barking deer with spotted deer group/herd. This association by solitary living barking deer can be for foraging effectively by reducing vigilance efforts in a group.Item The effect of disturbance-induced changes in vegetation structure and arthropod abundance on mixed-species bird flocks in the oak forests of the Western Himalaya(Wildlife Institute of India, Dehradun, 2024) Satish, Aditya; Page, Navendu; Bangal, Priti; Shahabuddin, GhazalaMixed-species flocks are an interactive group of insectivorous birds that forage together and move in the same direction. Flocks provide foraging and anti-predatory benefits to participants. Habitat disturbance may affect flocks by – a) altering presence or abundance of participating birds in the community or b) altering flocking propensities of species. 2. I investigated the effects of habitat disturbance on flocks in the Western Himalaya of Uttarakhand, India. Disturbance was mediated through two mechanisms – changes in vegetation structure and arthropod abundance (food availability). I examined how these environmental variations impacted flocks at a community level through changes in flock richness, flock size, number of flocks encountered and flock composition. I also explored how flocking propensity (a species-specific tendency to join flocks) is impacted by these variables. 3. Flocks are a subset of the insectivorous bird community. I found that insectivore richness and abundance decreased with reduced structural complexity of vegetation caused by disturbance. Reduced structural complexity also led to a corresponding decline in flock richness, size and number of flocks encountered in disturbed sites. With regards to variation of flock composition with disturbance, I found no clear species-composition-based clustering in transects with similar disturbance levels. 4. There seems to be an interactive link between arthropods, insectivorous birds and disturbance. Disturbance was correlated with strong declines in foliage arthropod abundance, which in turn contributes to lower insectivore richness and abundance. Flying arthropod abundance largely remained constant with disturbance, so I would like to draw 2 attention to the effect of foliage arthropod abundance on flock variables in the following sections. 5. The number of flocks encountered increased with foliage arthropod abundance. Since insectivores in undisturbed sites are supported by a larger prey base, the higher frequency of flocking may be correlated to the larger pool of insectivores that are available to flock at a given time. I did not find significant effects of foliage arthropod abundance on flock richness and flock size. 6. I found that flocking propensity of the three most common species (Gray-hooded Warbler, Black-throated Tit and Green-backed Tit) in flocks showed a declining trend with higher disturbance. I also observed that the propensity of these species showed an increasing trend with higher foliage arthropod abundance. However, none of these relationships were statistically significant. More detailed research is recommended to investigate these preliminary patterns further, to better understand the complex interactions between propensity, disturbance and food availability. 7. To conclude, disturbance was found to have a negative impact on mixed-species flocks, primarily by altering the presence and abundance of insectivorous birds. From a habitat perspective, insectivores suffered declines due to the simplification of vegetation structure. Insectivores also face food scarcity as foliage arthropod abundance decreases. The combined effect of habitat loss and reduced food availability in disturbed sites leads to lower insectivore richness and abundance, which is in turn reflected in fewer, smaller and more species-poor flocks.Item Nest site selection in Pallas’s Fish-eagle and nesting behaviour in the Rajaji-Corbett landscape(Wildlife Institute of India, Dehradun, 2024) Tangaria, Abhishek; Jha, Ashish; Pandav, Bivash; Kumar, R.S.This study investigates the nest site selection and nesting behavior of Pallas's Fish-eagle (PFE) across the sub-Himalayan tract in India, addressing a significant knowledge gap in the species' natural history. PFE, classified as Endangered globally and of high conservation concern nationally, exhibits unique nesting preferences and ecological requirements. Prior research, including studies from Bhutan and Bangladesh, has highlighted the species' preference for unobstructed nest trees and proximity to water sources and human settlements. However, comprehensive data from India remains limited. The study aims to explore factors influencing PFE's nest site selection, such as habitat quality, microclimate, food availability, and protection from predators and human disturbance. Additionally, it seeks to document the species' reproductive behavior, including hatchling development, parental duties, and nest predation pressures. Surveys covered segments of the Ganga, Kohlu, Kho, Palain, Mandal, Asan Barrage, Dakpatthar Barrage, and Ramganga, recording data on sightings and nest characteristics. A total of twelve nests were found, with only one destroyed by a storm. The majority of nests were built on Bombax ceiba and Shorea robusta trees, with an average height of 29.9m and GBH of 473cm. Behavioral observations were conducted at three accessible nests, monitoring parental presence, predator interactions, and nest guarding behavior. Spatial analysis in ArcGIS Pro considered various land cover types and environmental factors, finding that barren land and rangeland significantly influenced nest site selection. Euclidean distances to different land cover types and elevation were also key factors. Statistical analyses using R revealed correlations between tree height and nest height, and generalized linear models highlighted the impact of land cover and environmental variables on nest presence. Nest trees were typically tall and robust, with Bombax ceiba and Shorea robusta being the most commonly used species. These trees provided unobstructed branches and greater accessibility. The study also highlighted the importance of open land around the nest tree, aiding in vigilance and territory defense. At a macro scale, the presence of barren land and flooded regions positively influenced nest site selection, while slope and distance from rangeland showed negative correlations. Behavioral observations at three nests revealed high predation pressure, particularly from crows, and kites. The presence of human settlements increased predation incidents, affecting parental investment in nest defense.