WII Technical Reports/Books/Manuals
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Item Capacity building initiative on the dispersal and ranging patterns of elephants for effective management of human-elephant interactions(Wildlife Institute of India, Dehradun, 2022) Nigam, P.; Pandav, B.; Mondol, S.; Lakshmiarayanan, N.; Kumar, A.; Nandwanshi, V.B.; Das, J.; Biswas, S.; Udhayaraj, A.D.; Vishwakarma, R.; Habib, B.; Miachieo, K.; Narasingh Rao, P.V.Wild Asian elephant (Elephas maximus) populations are distributed in four major regions namely North West, North-East, East-Central and Southern regional meta-populations across India. Amongst them, the East-central regional population spread across the States of Odisha, Jharkhand, southern West Bengal, Chhattisgarh, and lately in Madhya Pradesh suffers disproportionately high levels of human elephant conflict. Among the myriad challenges facing management of human-elephant conflict in the region, elephant range expansion into new areas is overriding. One such range expansion that resulted in acute human-elephant conflict is being witnessed in the State of Chhattisgarh. Although northern Chhattisgarh was historically an elephant range, elephants reportedly disappeared during the period 1920 to late 1980s. While episodes of sporadic elephant occurrence in Chhattisgarh was reported during the period 1988- 1993, contemporary range expansion and concomitant human-elephant conflict began from the year 2000, and has accelerated during the last one decade. Faced with an enormous challenge of managing human-elephant conflict that is spatiotemporally dynamic unlike that of other elephant range States, constrained by limited Institutional capacities to assess and deal with the issue. Chhattisgarh Forest Department has been trying diversity its conflict mitigation strategies. Recognizing the need to objectively evaluate human-elephant conflict situation in the State, during the year 2017 Chhattisgarh Forest Department invited Wildlife Institute of India to conduct ecological research on elephants in Chhattisgarh with a three-year budget outlay. The project was a collaborative effort between Chhattisgarh Forest Department and WII. Considering the scope of the project, the project duration was further extended and eventually, the project lasted for the period July 2017 to March 2022. Being the final project report, the activities carried out as part of the project is summarized as under. Distribution and Demography In Chhattisgarh, the elephant distribution during the period 2012 to 2017 was reported from 16 Forest Divisions and four Protected Areas in the north and north-central regions of the state. The elephant population, as enumerated by Chhattisgarh Forest Department during 2021 , ranged from 250 to 300. The adult sex ratio recorded during the study was 1: 4.5. About 44% of the female segment of the population comprised of adults. Chhattisgarh elephant population is presently contiguous with other elephant populations in the neighboring states i.e., Madhya Pradesh, Jharkhand and in Odisha occurring as a meta-population 1 and thus cannot be considered as an isolated population. However, within Chhattisgarh, the population is relatively small and it occurs scattered over a large area as small and disjunctive groups facing a perpetual risk of getting isolated by ongoing linear infrastructure and other associated developmental activities in the State. If such groups get isolated, then they will not be viable in the long run. 1 Meta-population: Population of small populations that are connected through dispersals 1 O. ~ . -~ 1 -WU.d.U.fe .In.s-titu-te. o.f .In.di a Home Range, Movement Patterns & Dispersal, and Habitat Selection by Elephants During the period 2018-2022, WII-CGFD collaborative effort resulted in 10 elephant radio collaring in Chhattisgarh. The resultant effort provided 3106 elephant days of tracking information. Each of the radiocollared elephants provided an average of 310.6 (± 273) days of tracking data. As on 31 51 March 2022 when WII-CGFD collaborative project ended, two of the collared elephants (SD - Sehradev and MT - Maitri) were having functional collars. The estimated average home range (95% minimum convex polygon) of elephants in Chhattisgarh was 3172.8 km2 (± 2002.2 km2, Range: 462.3 - 6969.7 km2). The 95% kernel density home ranges of elephants were much lower averaging 512.3 km2 (± 235.3 km2, Range: 126.5 - 748.9 km2). The elephant home ranges were not wholly well defined, and marked by inter-annual shifts caused by exploratory behaviour. The elephant home ranges were relatively large. The dry season home ranges were significantly lower than monsoon and winter ranges. However, dry season home ranges of elephants are larger. The present study indicates that habitat quality in some of the forest patches - particularly those that are large and contiguous with minimal of human interference can potentially support elephants in the landscape. Thus, dry season ranges of elephants could serve as a surrogate for habitat quality. Monthly variations in home ranges were significant, and best explained by idiosyncrasies of individual elephants. Among the forest types open, moderately dense and very dense forests classified by Forest Survey of India based on crown densities, elephants selected open forests, that were predominantly juxtaposed with human-use areas. Although the crown density was low, the patches of open forests support dense stands of Sal (Shorea robusta) coppice with rank undergrowth offering adequate cover for elephants. Elephant habitat selection of these open forest patches appears to be influenced by potential foraging opportunities in human-use areas, and further facilitated by low inter-patch distance. Genetic Structure of Elephants Using 258 genetic samples collected from 9 Forest Divisions, elephant genetic structure in northern Chhattisgarh was evaluated. Analysis indicates that at least two different elephant lineages occur in Chhattisgarh. This implies that elephants occurring in Chhattisgarh have possibly come from different areas. Within the two different lineages, high relatedness amongst the individuals was observed corroborating with the general social structure of Asian elephant clans where individuals are mostly related. Crop Losses and Human Fatalities due to Elephants Crop losses caused by elephants were acute and widespread in Chhattisgarh. To draw an analogy, Karnataka's ex gratia payment towards crop losses by elephants during the period 2015-2020 was comparable with Chhattisgarh, although the former's elephant population is 93% more than the latter. The landscape-level assessment covering the whole of northern Chhattisgarh, and fine-scale assessment covering select areas in Surguja circle identified correlates of crop losses at both spatial scales. Elephant-related human deaths were widespread in the state. However, nearly 70% of incidences occurred in areas of high intensity of habitat-use by elephants. The human fatalities due to elephants were both temporally and spatially auto-correlated. 2Item PAN-India assessment and monitoring of endangered species covered under the Integrated Development of Wildlife Habitat Program (IDWH) - Snow leopard(Wildlife Institute of India, Dehradun, 2025) WII-MoEFCCIn 2019, the MoEF&CC launched the 'Snow Leopard Population Assessment in India (SPAI)* program, as part of India's active participation in the National and Global Snow Leopard and Ecosystem Protection Program (NSLEP & GSLEP). India has contributed in three significant landscapes to NSLEP & GSLER viz. Hemis-Spiti, Nanda Devi-Gangotri, and Khangchendzonga-Tawang. Alongside twelve National Snow Leopard and Ecosystem Priorities (NSLEPs), these efforts are aimed at conserving snow leopard and its habitats. The SPAI was designed to generate scientifically robust national and state-wise population estimates of the snow leopard across the high altitude habitats both inside and outside protected areas. Its overarching goal is to gather reliable data to guide effective conservation efforts and policy decisions. Based on the consultations with the Chief Wildlife Wardens (CWLWS) of the states and Union Territories (UTs) the SPAI sampling exercise was effectively executed across the PAN-INDIA Assessment and Monitoring of Endangered Species Covered Under the Integrated Development of Wildlife Habitats Program (IDWH) - SNOW LEOPARD Union Territories of Ladakh and Jammu & Kashmir and four snow leopard range states viz. Himachal Pradesh, Uttarakhand, Sikkim, and Arunachal Pradesh. The SPAI exercise was conducted from 2019 to 2023 following a two-step framework. Within this, the STEP-I involved a meticulous evaluation of snow leopard spatial distribution, incorporating habitat covariates into the analysis. This approach, founded on occupancy modeling, relied on data sourced from sign and interview surveys, comprehensive fieldwork, and precise geospatial mapping. This assessment culminated in the refinement of the snow leopard distribution map and the establishment of a foundational dataset for the stratification of snow leopard population sampling. In the STEP- II, snow leopard abundance estimation was achieved through camera trapping. Furthermore, comprehensive abundance assessments were conducted for the primary prey species at selected sites. Given the constraints posed by the situations of Covid-19, some states such as Sikkim and Arunachal Pradesh conducted the STEP-I and II exercises simultaneously. Till 2016, a substantial third of the snow leopard range (spanning ca. 100,347 km2) had not received research attention, which was reduced to just 5%, mainly in small pockets of Ladakh, Jammu & Kashmir; Uttarakhand, and Himachal Pradesh. However; status surveys in the recent years have increased the understanding on preliminary information about snow leopard or prey is available for 80% of the snow leopard range (ca. 79,745 km2), compared to 56% in 2016. To provide robust information on the snow leopard numbers, the SPAI exercise involved surveying the snow leopard habitats using a substantial network of camera traps. In total, the SPAI sampling exercise utilized 1,971 camera trap locations, leading to the identification of 241 unique individuals. These comprehensive findings allow for the estimation of a total snow leopard population of 718 individuals in India. A total of 126 unique snow leopard individuals were identified from the Union Territory of Ladakh, across a total of 956 camera trap locations, yielding an estimated density of 0.34 individuals/100 km2 and a population of477 individuals. Camera trapping efforts in 135 sites in the Union Territory of Jammu & Kashmir led to detection of 9 unique individuals across the snow leopard range. The estimated density was 0.75 individuals/100 km2 and population of 9 individuals. For the snow leopard range states, Himachal Pradesh recorded a total of 44 unique individuals identified across 284 camera trap locations. The density of snow leopard in the state ranged from 0.08 to 0.37 individuals/100 km2. The exercise was indicative of a mean population size of 51 individuals of snow leopard in Himachal Pradesh. The state of I PAN-INDIA Assessment and Monitoring of Endangered Species Covered Under the Integrated Development of Wildlife Habitats Program (IDWH) - SNOW LEOPARD Uttarakhand recorded a total of 40 unique individuals, with a density ranging from 0.7 to 1.04 individuals/100 km2. The population estimates of snow leopard in Uttarakhand was 124. In the eastern Himalayan states, Sikkim, with a total of 99 camera trap locations, recorded a total of 14 unique individuals with an estimated density of 0.40 snow leopards/100 km2. The estimate was 21 snow leopards for this state. On the other hand, in Arunachal Pradesh, 8 unique snow leopards were identified across 115 camera trap locations. The density was 0.26 individuals/100 km2 and the population was estimated as 36 individuals in the state. In the Indian Himalayan region, two prey species dominate the diet of the snow leopard: the bharal or blue sheep (Pseudois nayaur), and the Siberian ibex (Capra sibirica). Bharal exhibit a near-total range overlap with snow leopard across the greater and trans-Himalayan regions, while ibex occupy extensive ranges within the northwestern trans-Himalayan region. In the subalpine and alpine region Himalayan tahr and musk deer form important component to snow leopard diet. Other mountain ungulates, though less prominent, contribute to the diet and hold local importance. Himachal Pradesh is the only snow leopard range state in India where systematic surveys on its primary prey species have been conducted. Conversely, states like Arunachal Pradesh and Jammu & Kashmir remain significantly understudied, while data from Uttarakhand and Sikkim are fragmented and localized. Ladakh, due to its visibility and accessibility, has a relatively well-documented history of ungulate monitoring. Limited long term data suggest a decline in ungulate populations across the Himalaya, except for the Tibetan antelope (Pantholops hodgsoni), which has shown signs of recovery. Markhor (Capra falconeri) habitat shrank from 300 km2 in the 1940s to just 120 km2 by 2004-2005. Similarly, musk deer (Moschus spp.) in Kedarnath Wildlife Sanctuary (Uttarakhand) have seen a steady decline from 3.7 individuals/km2 (1989-1991) to 1.2 individuals/km2 by 2000. A similar declining trend is evident in ibex (Capra sibirica) populations in Pin Valley National Park, Himachal Pradesh. Density estimates dropped from 2.3 individuals/km2 in 1989 to 1.2 individuals/km2 in 1997, with a further reduction to 0.37 individuals/km2 by 2012. These declines underscore the urgent need for targeted conservation efforts and systematic prey monitoring to ensure the long-term viability of snow leopard populations.Item Ecology of Dhole (Cuon alpinus Pallas) in Central India(Wildlife Institute of India, Dehradun, 2006) Acharya, Bhaskar B.; Johnsingh, A.J.T.; Sankar, K.The objectives of the project in Pench Tiger Reserve, Madhya Pradesh, were to estimate the seasonal abundance of dhole prey species, the diet of the dhole packs from their scats and kills, to determine patterns of habitat use and range sizes of dhole packs, to estimate temporal changes in size and composition of dhole packs, and the processes behind such changes, to screen captured dhole and other animals for diseases, and estimate the probability of contact with potential disease carriers, to devise standardised protocols for dhole population surveys and to estimate dhole population size for the Tiger Reserve.Item Long term conservation plan for hangul Part II Hangul movement pattern study using GPS satellite telemetry - final report(2016-20)(Wildlife Institute of India, Dehradun, 2021) Ahmad K.The globally viable single population of Kashmir Red Deer or Hangul (Cervus hanglu hanglu), one of the critically endangered subspecies of the Central Asian Deer is restricted to a confined area of 141 km2 Dachigam National Park (34°05ʹ to 34°32ʹN; 74°50ʹ to 75°16ʹE) in the Greater Himalayan mountain range of the Northwest Himalayan biogeographic region zone 2A, with some stray populations occurring in the adjoining relic range areas (Ahmad et al. 2009; Qureshi et al. 2009). Earlier, the species was widely distributed in the mountain of Kashmir Himalayas along the entire Greater Himalayan mountain range (Gee 1965; Schaller 1969; Prater, 1993; Nowak, 1999) declined drastically in the recent past from 5000 Individuals prior to 1947 to less than 200 Individuals at present. Current trends in the Hangul population indicate that the species could go extinct if necessary serious interventions are not made immediately and as such there was need to undertake urgent measures to hold the declining trends in the Hangul population (Ahmad et.al. 2009: Qureshi et.al. 2009; Ahmad et.al 2013). Therefore, understanding the ecology and biology particularly the movement ecology of this critically endangered deer species with small single population was fundamental to develop better strategies for conservation and management practices. The present study duly funded by the MOEF & CC, Government of India was as such initiated to understand the lesser known aspects of movement ecology and behaviour of this last viable population of the Hangul for its effective management, conservation planning and species population recovery under the following objectives: 1) Studying the seasonal Home range size and ranging and movement patterns of Hangul in and outside Dachigam National Park vis-à-vis Hangul migration route, important stop-over sites, and barriers and corridors to migration into the Hangul’s relic areas.2) Studying the lesser known aspects of Hangul ecology viz., habitat use, activity patterns, behaviour and predation prerequisite for effective long term management and conservation of Hangul and its habitats. 3) To identify the potential habitats used by Hangul outside Dachigam and assess and evaluate the extent and magnitude of habitat conditions and threats therein. 4) To identify threats, anthropogenic pressures and other factors particularly predation pressure by leopard and meso-carnivores that impact Hangul distribution and movement patterns. The capture and Satellite collaring of five Hangul (2 males, 3 female) successfully conducted for the first time under this project has been a milestone achievement in the field of satellite telemetry. The findings of this research study indicated that the Home range size varied from 4.98 Km2 in spring to 7.83 Km2 in summer. One of the female collared Hangul showed movement patterns outside Dachigam National Park towards Sindh forest division crossing the river Sindh and covering an area of 137.94 km, with area use of 10.86-12.26 Km2 in summer 2019 to 137.94 km in summer 2020 and a maximum home range of 124.4 km2 in Summer 2020 to colonize and establish its new summer habitat in the erstwhile range area of Wangath-Naranag Conservation Reserve (CR). The data and information generated has enabled us to identify the corridor areas of movement of the Hangul from Dachigam National Park and outside in the 3 Long Term Conservation Plan for Hangul Part II: Hangul Movement Pattern Study Using GPS-Satellite Telemetry adjoining erstwhile range areas in north and south and habitats assessment therein. The data generated also indicated that Hangul shows two activity peaks in morning and evening hours with significant seasonal variations. The findings of the study are of great ecological significance as the significant information generated through this research on the lesser known aspects of movement ecology including animal home ranges and habitat use, biology and behaviour of the Hangul deer would go long way in supporting the management interventions for population recovery and long term survival of this endemic deer of India in Dachigam National Park and its erstwhile range areas in Kashmir Himalayas. Major management and conservation Intervention recommendations 1. The study revealed that major and viable population of Hangul are confined to Dachigam National Park. Despite availability of ideal summer habitats for the Hangul in upper Dachigam, these alpine meadow habitats are not being explored or used by Hangul. The satellite collared Hangul movements indicated that the animals showed upward movements to Dagwan alpine meadows of upper Dachigam but restricted their movements further in to the alpine meadows, possibly due to heavy disturbances of excessive livestock grazing there. 2. Management interventions are as such required towards expansion of Range of Hangul to alpine meadows of Upper Dachigam and potential corridor areas outside Dachigam NP identified through this research, so that these ideal summer habitats are recuperated and used by Hangul in summer as it used to in the past and to ensure gene flow between the Dachigam and adjoining range populations. 3. Hangul conservation breeding-cum- reintroduction programme is imperative to expand the range of Hangul by restocking and augmenting the small isolated Hangul populations in its relic range areas outside Dachigam National Park starting with the Overa Wildlife Sanctuary which has ideal disturbance free habitats available. 4. The Hangul species recovery programme through a project mode by initiation of Project Hangul on the pattern of Project Tiger is crucial to ensure Hangul species recovery and long term survival of the species and its landscapes in the region. 5. This research study as indicated by earlier studies by the Investigator (s) has revealed that besides poaching and continued degradation of Hangul summer habitats in Upper Dachigam, along with biotic interference in winter habitats, low breeding, female biased sex ratio, the problem of survival of the young and inadequate recruitment of calf to adulthood due to factors such as considerable predation by common Leopard, Asiatic Black Bear, dogs and meso-carnivores (Fox and Jackal) are major challenges for the long term survival of the Hangul in the landscape. 6. The study revealed a significant contribution of Hangul in the diet of Golden jackal (9.09%) and Red fox (6.45%). These ecological issues threatening the long term survival of Hangul need to be investigated and addressed further on long term basis through initiating a breeding biology study to better understand the causes of low breeding and fawn/calf survival in the Hangul population in Dachigam National Park and the adjoining landscape. 7. This research study and the earlier studies by the Investigator (s) has indicated that species due to its small population size, restricted range distribution, critically endangered status, ecological threats long Term Conservation Plan for Hangul Part II: Hangul Movement Pattern Study Using GPS-Satellite Telemetry and potentially low genetic variation is at the brink of extinction and needs immediate management interventions to reverse the declining trend in the population. The regulated monitoring of the Hangul populations on a long-term scientific basis using latest techniques of satellite collaring, camera trapping and population genomics is imperative. 8. Strengthening Hangul genome sequencing to understand the DNA mitochondrial based phylogeography of the species and Skull based genetic investigations to link the mitochondrial DNA analysis findings with the nuclear genetic analysis to further establish the degree of closeness or divergence between Hangul and the Bactrian deer.